რაოდენობრივი ეკოლოგიის პროგრამა 

2008-2020 წწ.

1. Tarkhnishvili, D., Gabelaia, M., & Adriaens, D. (2020). Phenotypic divergence, convergence and evolution of Caucasian rock lizards (Darevskia). Biological Journal of the Linnean Society, 130(1), 142-155.
2. Tarkhnishvili, D., Barateli, N., Murtskhvaladze, M., & Iankoshvili, G. (2020). Estimating phenotypic heritability of sexual and unisexually reproducing rock lizards (genus Darevskia). Zoologischer Anzeiger, 285, 105-113.
3. Murtskhvaladze, M., Tarkhnishvili, D., Anderson, C. L., & Kotorashvili, A. (2020). Phylogeny of caucasian rock lizards (Darevskia) and other true lizards based on mitogenome analysis: Optimisation of the algorithms and gene selection. Plos one, 15(6), e0233680.
4. Tarkhnishvili, D., Yanchukov, A., Şahin, M. K., Gabelaia, M., Murtskhvaladze, M., Candan, K., ... & Ilgaz, Ç. (2020). Genotypic similarities among the parthenogenetic Darevskia rock lizards with different hybrid origins. BMC Evolutionary Biology, 20(1), 1-25.
5. Galoyan, E., Moskalenko, V., Gabelaia, M., Tarkhnishvili, D., Spangenberg, V., Chamkina, A., & Arakelyan, M. (2020). Syntopy of two species of rock lizards (Darevskia raddei and Darevskia portschinskii) may not lead to hybridization between them. Zoologischer Anzeiger, 288, 43-52.
6. Shakarashvili, M., Kopaliani, N., Gurielidze, Z., Dekanoidze, D., Ninua, L., & Tarkhnishvili, D. (2020). Population genetic structure and dispersal patterns of grey wolfs (Canis lupus) and golden jackals (Canis aureus) in Georgia, the Caucasus. Journal of Zoology, 312(4), 227-238.
7. Zafar-ul Islam, M., Volmer, R., al Boug, A., Shehri, A. A., & Gavashelishvili, A. (2020). Modelling the effect of competition for prey and poaching on the population of the Arabian Leopard, Panthera pardus nimr, in Saudi Arabia (Mammalia: Felidae). Zoology in the Middle East, 66(2), 95-106.
8. Islam, M. Z., Gavashelishvili, A., Kokiashvili, L., al Boug, A., as Shehri, A., Peterson, A. T., & García, D. J. (2020). Modeling the distribution and movement intensity of Arabian Leopard Panthera pardus nimr in the Arabian Peninsula. Authorea Preprints.
9. Kuemmerle, T., Bluhm, H., Ghoddousi, A., Arakelyan, M., Askerov, E., Bleyhl, B., Gavashelishvili, A., ... & Manvelyan, K. (2020). Identifying priority areas for restoring mountain ungulates in the Caucasus ecoregion. Conservation Science and Practice, 2(11), e276.
10. Mikeladze, G., Gavashelishvili, A., Akobia, I., & Metreveli, V. (2020). Estimation of forest cover change using Sentinel-2 multi-spectral imagery in Georgia (the Caucasus). iForest-Biogeosciences and Forestry, 13(4), 329.
11. Chaladze, G. (2020). Estimation of Ground Dwelling Arthropod Densities Using Pitfall Traps: Individual Based Simulations. Evolutionary Biology, 5(1), 1-5.
12. Zerbe, S., Pieretti, L., Elsen, S., Asanidze, Z., Asanidze, I., & Mumladze, L. (2020). Forest Restoration Potential in a Deforested Mountain Area: An Ecosociological Approach towards Sustainability. Forest Science, 66(3), 326-336.
13. Mumladze, L., Japoshvili, B., & Anderson, E. P. (2020). Faunal biodiversity research in the Republic of Georgia: a short review of trends, gaps, and needs in the Caucasus biodiversity hotspot. Biologia, 75(9), 1385-1397.
14. Thormann J, Ahrens D, Anderson C, Astrin JJ, Mumladze L , Rulik B, Tarkhnishvili D, Espeland M, Geiger M, Hein N, Iankoshvili G, Karalashvili E, Mengual X, Morkel C, Neiber MT, Peters RS, Reimann A, Ssymank A , Wesener T, Ziegler J, Misof B. 2019. A prelude to the Caucasus Barcode of Life Platform (CaBOL): Biodiversity Days in Georgia in 2018 and 2019. Bonn zoological Bulletin 68 (2): 275–296.
15. Barjadze S., Asanidze Z., Gavashelishvili A., Soto-Adams FN. 2019. The hypogean invertebrate fauna of Georgia (Caucasus). Zoology in the Middel East 65, 1-10.
16. Bleyhl B, Arakelyan M, Askerov E, Bluhm H, Gavashelishvili A, Ghasabian M, Ghoddousi A, Heidelberg A, Khorozyan I, Malkhasyan A, Manvelyan K, Masoud M, Moqanaki EM, Radeloff VC, Soofi M, Weinberg P, Zazanashvili N, Kuemmerle T. 2019. Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Diversity and Distributions 25: 129-141.
17. H Pretzsch, M Steckel, M Heym, P Biber, C Ammer, M Ehbrecht, K Bielak, F Bravo, C Ordóñez, C Collet, F Vast, L Drössler, G Brazaitis, K Godvod, A Jansons, J de-Dios-García, M Löf, J Aldea, N Korboulewsky, DOJ Reventlow, A Nothdurft, M Engel, M Pach, J Skrzyszewski, M Pardos, Quentin Ponette, R Sitko, M Fabrika, M Svoboda, J Černý, B Wolff, R Ruíz-Peinado, M del Río. 2019. Stand growth and structure of mixed-species and monospecific stands of Scots pine (Pinus sylvestris L.) and oak (Q. robur L., Quercus petraea (MATT.) LIEBL.) analysed along a productivity gradient through Europe. European Journal of Forest Research 
18. Géraud de Streel, C Collet, I Barbeito, K Bielak, Andres Bravo-Oviedo, G Brazaitis, L Coll, L Drössler, D Forrester, M Heym, M Löf, M Pach, H Pretzsch, R Ruiz-Peinado, J Skrzyszewski, J Stankevičiūtė, M Svoboda, K Verheyen, T Zlatanov, D Bonal, Quentin Ponette. 2019. Contrasting patterns of tree species mixture effects on wood δ13C along an environmental gradient. European Journal of Forest Research. 
19. Peter Freer-Smith, Bart Muys, Michele Bozzano, Lars Drössler, Niall Farrelly, Hervé Jactel, Jaana Korhonen, Gianfranco Minotta, Maria Nijnik, Christophe Orazio. 2019. Plantation forests in Europe: challenges and opportunities. European Forest Institute 9, 1-52.
20. Gabelaia M, D Tarkhnishvili, D Adriaens. 2018. Use of three-dimensional geometric morphometrics for the identification of closely related species of Caucasian rock lizards (Lacertidae: Darevskia). Biological Journal of the Linnean Society, Volume 125, Issue 4, 26 November 2018, Pages 709–717,
21. Gavashelishvili A., YA Yarovenko, EA Babayev, GMikeladze, Z Gurielidze, D Dekanoidze, N Kerdikoshvili, L Ninua, N Paposhvili. 2018. Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus. Journal of Mammalogy, xx(x):1–13,
22. Axel Barlow, James A Cahill, Stefanie Hartmann, Christoph Theunert, Georgios Xenikoudakis, Gloria G Fortes, Johanna LA Paijmans, Gernot Rabeder, Christine Frischauf, Aurora Grandal-d’Anglade, Ana García-Vázquez, M Murtskhvaladze, Urmas Saarma, Peeter Anijalg, Tomaž Skrbinšek, Giorgio Bertorelle, Boris Gasparian, Guy Bar-Oz, Ron Pinhasi, Montgomery Slatkin, Love Dalén, Beth Shapiro, Michael Hofreiter, 2018. Partial genomic survival of cave bears in living brown bears. Nature ecology & evolution 2: 1563
23. M Nassirkhani, L Mumladze 2018. A new cave pseudoscorpion (Pseudoscorpiones: Neobisiidae) from western Georgia. Arachnology 17 (9), 496-500
24. M Murvanidze, N Todria, L Mumladze, L Kalatozishvili. 2018. Diversity of soil mite communities in different habitats of Saskhori quarries, Georgia. Persian Journal of Acarology 7
25. Sophio Gabelashvili, Levan Mumladze, Ani Bikashvili, Pavel Sroka, Roman J Godunko, Bella Japoshvili. 2018. The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country. Turkish Journal of Zoology 42: 252-262.
26. T Kuljanishvili, L Mumladze, L Kalous, B Japoshvili 2018. Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia). Biologia 73 (1), 93-100
27. Ninua L, Tarkhnishvili D, Gvazava E. 2018. Phylogeography and taxonomic status of trout and salmon from the Ponto ‐Caspian drainages, with inferences on European Brown Trout evolution and taxonomy. – Ecology and Evolution 8: 2645-2658
28. Gabelashvili A, Mumladze M, Bikashvili A, Sroka P, Godunko RJ, Japoshvili B. 2018. The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country. Turkish J. Zool. 42: 252-262.
29. Kuljanishvili T, Mumladze L, Kalous L, Japoshvili B. 2018. Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia). Biologia, 73: 93-100.
30. Tarkhnishvili D., M Gabelaia, A Kandaurov, A Bukhnikashvili, G Iankoshvili. 2017. Isolated population of the Middle Eastern Phoenicolacerta laevis from the Georgian Black Sea Coast, and its genetic closeness to populations from southern Turkey. Zoology in the Middle East. 63, 311-315
31. Gabelaia M, D Adriaens, D Tarkhnishvili. 2017. Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger-A Journal of Comparative Zoology 268, 32-40
32. Tarkhnishvili D, M Murtskhvaladze, Cl Anderson. 2017. Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linenan Society, 121, 365–378.
33. Tarkhnishvili D, A Gavashelishvili, M Murtskhvaladze, A Latsuzbaia. 2017. Landscape Complexity in the Caucasus impedes Genetic Assimilation of Human Populations More Effectively than Language or Ethnicity. Human Biology 88, 287-300
34. Mumladze L, W Ulrich, Z Asanidze, G Japoshvili. 2017. An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Écoscience, 1-5
35. Mumladze L, M Murvanidze, M Maraun. 2017. Patterns of oribatid mite species diversity: testing the effects of elevation, area and sampling effort. Experimental and Applied Acarology, 1-18.
36. Aslan E, L Mumladze, G Japoshvili. 2017. List of leaf beetles (Coleoptera: Chrysomelidae) from Lagodekhi reserve with new records for Transcaucasia and Georgia, Zootaxa 42277 (1), 086-098
37. Japoshvili B, L Mumladze, L Murvanidze. 2017. The population of Carassius gibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iranian Journal of Fisheries Sciences 16 (2), 793-799
38. Mumladze L, Z Asanidze, F Walther, B Hausdorf. 2017. Beyond elevation: testing the climatic variability hypothesis vs. Rapoport’s rule in vascular plant and snail species in the Caucasus. Biological Journal of the Linnean Society, 121, 753–763
39. Chaladze G. 2017. Taxonomy should be more photography based-eliminate need of physical specimen to study morphology. Zootaxa 4247 (3), 331-331
40. Chaladze G, L Kalatozishvili, Z Janiashvili, G Bakuradze. 2017. Pterostichus neilgaimani sp. nov., a new species of ground beetles (Coleoptera: Carabidae) from a relict sacred grove in Eastern Georgia. Zootaxa 4329 (1), 91-95.
41. Gavashelishvili A, D Tarkhnishvili. 2016. Biomes and human distribution during the last ice age. Global Ecology and Biogeography, DOI: 10.1111/geb.12437 ISI IF 5.84
42. Mumladze, L., Asanidze, Z., Walther, F., Hausdorf, B. (2017). Beyond elevation: Testing the climatic variability hypothesis versus Rapoport’s rule in vascular plant and snail species in the Caucasus. Oecologia (Accepted). ISI IF 2.902
43. Chaladze G. 2016. Heterosexual Male Carriers Could Explain Persistence of Homosexuality in Men: Individual-Based Simulations of an X-Linked Inheritance Model. Archives of Sexual Behavior, 1-7. ISI IF 2.704
44. Tarkhnishvili D, M. Murtskhvaladze, CL Anderson. 2017. Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linnean Society (Accepted) ISI IF 1.984
45. Murvanidze M, L Mumladze. 2016. Annotated checklist of Georgian oribatid mites. Zootaxa 4089 (1), 1-81. ISI IF 0.994
46. Tarkhnishvili D, M Gabelaia, L Mumladze, M Murtskhvaladze. 2016. Mitochondrial phylogeny of the Darevskia saxicola complex: two highly deviant evolutionary lineages from the easternmost part of the range. he Herpetological Journal 26 (2), 175-182. ISI IF 0.808
47. Tarkhnishvili D, A. Gavashelishvili, M Murtskhvaladze (2017) Landscape Complexity in the Caucasus Impedes Genetic Assimilation of Human Populations More Effectively Than Language or Ethnicity. Human Biol. (Accepted). ISI IF 0.857
48. Japoshvili, B., Mumladze, L., Murvanidze, L. (2017). The population of Carssiusgibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iran J Fish Sci (Accepted). ISI IF 0.393
49. Murvanidze M, L Mumladze, T Arabuli, S Barjadze, M Salakaia. 2016. Oribatida diversity in different microhabitats of Mtirala National Park. J. Acarol. Soc. Jpn 25 (Supplement1), S35-S49. ISI IF N/A
50. Bakhtadze, N.G., Chakvetadze, N.L., Mumladze, L.J., Bakhtadze, G.I., & Tskhadaia, E.A., (2016). Karyological Data of Terrestrial Mollusks (Mollusca: Gastropoda: Pulmonata) of Georgia. Proceedings of the Institute of Zoology 12/2016; XXV: 23-27 ISI IF N/A
51. Gabelashvili, S., Bikashvili, A., Shubitidze, Z., Gioshvili, M., Pankvelashvili, E., Mumladze, L., & Japoshvili, B. (2016). Family level diversity and distribution of macroinvertebrates of Madatapa, Khanchali and Bughdasheni lakes in Javakheti plateau (South Georgia). Proceedings of the Institute of Zoology 12/2016; XXV: 117-127 ISI IF N/A
52. Mumladze, L., & Paposhvili, N. (2016). A New Addition to the Malacofauna of Georgia – Eobania Vermiculata is Replenishing its Range. Proceedings of the Institute of Zoology 12/2016; XXV:153-155 ISI IF N/A
53. Seehausen, M., Schröter, A., Mumladze, L., Grebe, B. (2016) Additional Odonata records from Georgia, southern Caucasus ecoregion,with the first record of Ischnura fountaineae (Odonata:Coenagrionidae). Notulae Odonatologicae. 8 (8), 266-283 ISI IF N/A
54. Gabelaia, M., Tarkhnishvili, D. and Murtskhvaladze, M. 2015. Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta. Amphibia-Reptilia 36: 45-54.
55. Mumladze, L., Murvanidze, M., Maraun, M. & Salakaia, M. (2015). Oribatid mite communities along an elevational gradient in Sairme gorge (Caucasus). Experimental and Applied Acarology. 66(1), 41-51.
56. Farhadinia, M. S., Farahmand, H., Gavashelishvili, A., Kaboli, M., Karami, M., Khalili, B., Montazamy, S. 2015. Molecular and craniological analysis of leopard, Panthera pardus (Carnivora: Felidae) in Iran: support for a monophyletic clade in Western Asia. Biological Journal of the Linnean Society 114: 721-736.
57. Lewis NS, Verhagen JH, Javakhishvili Z, Russell CA, Lexmond, P. et al (20 authors). 2015. Influenza A virus evolution and spatio-temporal dynamics in Eurasian Wild Birds: A phylogenetic and phylogeographic study of whole-genome sequence data. Journal of General Virology
58. Tarkhnishvili D. 2014. Rana macrocnemis. In: Grossenbacher, K. (ed.), Handbuch Amphibien & Reptilien Europas. Aula-Verlag, Germany.
59. Kopaliani, N., Shakarashvili, M., Gurielidze, Z., Qurkhuli, T. & Tarkhnishvili, D. 2014. Gene flow between wolves and shepherd dogs in the Caucasus. J. Hered. (in press).
60. Mumladze L, Cameron RAD, Pokryszko BM. 2014. Endemic land molluscs in Georgia (Caucasus): how well are they protected by existing reserves and national parks? Journal of Molluscan Studies 80: 67-73.
61. Tarkhnishvili D, Murtskhvaladze M, Gavashelishvili A. 2013. Speciation in Caucasian lizards: climatic dissimilarity of the habitats is more important than isolation time. Biological Journal of the Linnean Society, 109, 876-892.
62. Mumladze L., Tarkhnishvili D., Murtskhvaladze M. 2013. Systematics and evolutionary history of large endemic snails from the Caucasus (Helix buchii and H. goderdziana) (Helicidae) American Malacological Bulletin 31 (2), 225-234. 
63. Mumladze L, Murvanidze M, Behan-Pelletier V. 2013. Compositional patterns in Holarcticpeat bog inhabiting oribatid mite (Acari: Oribatida) communities Pedobiologia 55 (1), 41-48
64. Murvanidze M, Mumladze L, Arabuli T, Kvavadze E. 2013. Oribatid mite colonization of sand and manganeze tailing sites. Acarologia 53 (2), 203-215
65. Japoshvili B, Mumladze L, Kucuk F. 2013. Invasive Carassius Carp in Georgia: Current state of knowledge and future perspectives. Current Zoology, in press
66. Mumladze L. 2013. Shell size differences in Helix lucorum Linnaeus, 1758 (Mollusca: Gastropoda) between natural and urban environments. Turk J. Zool. 36: 1-7
67. Lewis NS , Z Javakhishvili, CA Russell, A Machablishvili, P Lexmond, JH ..2013.Avian Influenza Virus Surveillance in Wild Birds in Georgia: 2009–2011. PloS one 8 (3), e58534 2008-2012: 20
68. Tarkhnishvili, D. 2012: Evolutionary History, Habitats, Diversification, and Speciation in Caucasian Rock Lizards. Advances in Zoology Research. Owen P. Jenkins, Ed. 2, 79-120. Nova Scientific Publishers, NY.
69. Tarkhnishvili D., Gavashelishvili A., Mumladze L. 2012: Palaeoclimatic models help to understand current distribution of Caucasian forest species. Biological Journal of the Linnean Society. 105: 231-248.
70. Khorozyan I, Malkhasyan A, Murtskhvaladze M. 2012. The striped hyaena Hyaena hyaena (Hyaenidae, Carnivora) rediscovered in Armenia. Folia Zoologica 60: 253-261.
71. Murtskhvaladze M., Gurielidze Z., Kopaliani N., Tarkhnishvili D. 2012: Gene introgression between Gazella subguturrosa and G. marica: limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica 
72. Gavashelishvili A, McGrady M, Ghasabian M, Bildstein KL. 2012. Movements and habitat use by immature Cinereous Vultures (Aegypius monachus) from the Caucasus. Bird Study 59 (4): 449-462.
73. Chaladze G. 2012: Climate-based model of spatial pattern of the species richness of ants in Georgia. Journal of Insect Conservation. 
74. Murvanidze M, Mumladze L, Arabuli T, Kvavadze E. 2011. Landscape distribution of oribatid mites (Acari, Oribatida) in Kolkheti National Park (Georgia, Caucasus). Zoosymposia 6: 221-233.
75. Murvanidze M, Kvavadze E, Mumladze L, Arabuli T. 2011. Comparison of Earthworms (Lumbricidae) and Oribatid Mite (Acari, Oribatida) Communities in Natural and Urban Ecosystems. Vestnik Zoologii, 45: 16-24.
76. Pokryszko B, Cameron R, Mumladze L, Tarkhnishvili D. 2011: Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society. 102: 239-250.
77. Tarkhnishvili, D. Chaladze, G. et al., 2011. Georgian biodiversity database. www.biodiversity-georgia.net  
78. Tarkhnishvili D., Gavashelishvili A., Avaliani A., Murtskhvaladze M., Mumladze L. 2010: Unisexual rock lizard might be outcompeting its bisexual progenitors in the Caucasus. Biological Journal of the Linnean Society. 101: 447-460.
79. Murtskhvaladze M., Gavashelishvili A., Tarkhnishvili D. 2010: Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus. Molecular Ecology. 19: 1829-1841.
80. Gavashelishvili A, Javakhishvili Z. 2009: Combining Radio-Telemetry and Random Observations to Model the Habitat of Caucasian Grouse (Tetrao Mlokosiewiczi). Oryx. 44: 491-500. 
81. Tarkhnishvili D., Serbinova I., Gavashelishvili A. 2009: Modelling the range of Syrian spadefoot toad (Pelobates syriacus) with combination of GIS-based approaches. Amphibia-Reptilia. 30: 401-412
82. Sayim, F., Baskale, E., Tarkhnishvili, D. Kaya, U. 2009. Some water chemistry parameters of breeding habitats of Caucasian salamander, Mertensiella caucasica in the Western Lesser Caucasus. Comptes Rendus Biologies, Vol. 332, No. 5. (May 2009), pp. 464-469.
83. Tarkhnishvili D., Kaya U. 2009: Status and Conservation of the Caucasian Salamander (Mertensiella caucasica). In: Zazanashvili, N., Mallon, D. (eds), Status And Protection of Globally Threatened Species in The Caucasus, pp. 157-165.
84. Gavashelishvili A. 2009: GIS-based Habitat Modeling of Mountain Ungulate species in the Caucasus Hotspot. In: Zazanashvili, N., Mallon, D. (eds), Status And Protection of Globally Threatened Species in The Caucasus, pp. 74-82
85. Sinsch U., Schneider H. & Tarkhnishvili D. 2008 Bufo bufo Superspecies – Erdkröten-Artenkreis. PP. 191-335 In: Grossenbacher, K. (ed.), Handbuch Amphibien & Reptilien Europas. Aula-Verlag, Germany
86. Mumladze L., Tarkhnishvili D., Pokryszko B. 2008: A new species of the genus Helix from south-western Georgia. Journal of Conchology 39: 483-485. 
87. Kvavadze E, Rukhadze L, Nikolaishvili V, Mumladze L. 2008. Botanical and zoological remains from an early medieval grave at Tsitsamuri, Georgia. Vegetation History and Archaeobotany 17 (1), 217-224
88. Angus RB, Galián J, Wrase DW, Chaladze G. 2008. The western Palaearctic species of the Pterostichus nigrita (Paykull) complex, with the description of a new species from Spain and a new subspecies of P. nigrita from Anatolia (Coleoptera, Carabidae). Nouv Rev Entomol 25, 297-316
89. Gavashelishvili A., Lukarevsky V. 2008: Modelling the habitat requirements of the leopard Panthera pardus in West and Central Asia. Journal of Applied Ecology 45: 579-588.
90. Poulakakis N, Antoniou A, Mantziou G, Parmakelis A, Skartsi T, Vasilakis D, Elorriaga J, De La Puente J, Gavashelishvili A, Ghasabyan M, Katzner T, Mcgrady M, Batbayar N, Fuller M, Natsagdorj T, 2008. Population structure, diversity, and phylogeography in the near‐threatened Eurasian black vultures Aegypius monachus (Falconiformes; Accipitridae) in Europe: insights from microsatellite and mitochondrial DNA variation. Biological Journal of the Linnean Society, 95: 859-872.
91. Tarkhnishvili D., Kaya U., Gavashelishvili A., Serbinova I. 2008: Ecological Divergence between two Evolutionary Lineages of the Caucasian Salamander: Evidence from the GIS analysis. Herpetological Journal. 18: 155-163

მსხვილი ძუძუმწოვრების ეკოლოგიის პროგრამა 

2006-2020 წწ.

1. Shakarashvili N. Kopaliani Z. Gurielidze  D. Dekanoidze  L. Ninua  D. Tarkhnishvil. 2020.  Population genetic structure and dispersal patterns of grey wolf (Canis lupus) and golden jackal (Canis aureus) in Georgia, the Caucasus. Journal of Zoology.
2. ნათია კოპალიანი, ზურაბ გურიელიძე, დავით დეკანოიძე, ივანე დარასელია, ნანა დევიძე. 2020. მართვის გეგმა ზურმუხტის ქსელის ტერიტორიისთვის სვანეთი 1 (Svaneti 1 – GE0000012). ილიას სახელმწიფო უნივერსიტეტი. მომზადდა გარემოს დაცვის და სოფლის მეურნეობის სამინისტროს ხელშეწყობით. თბილისი,2020
3. ნათია კოპალიანი, ზურაბ გურიელიძე, დავით დეკანოიძე, ივანე დარასელია, ნანა დევიძე. 2020. მართვის გეგმა ზურმუხტის ქსელის ტერიტორიისთვის სვანეთი 2 (Svaneti 2 – GE0000045). ილიას სახელმწიფო უნივერსიტეტი. მომზადდა გარემოს დაცვის და სოფლის მეურნეობის სამინისტროს ხელშეწყობით. თბილისი,2020
4. Pilot M, Moura A., Okhlopkov I, Mamaev N, Alagaili A., Mohammed O., Eduard Yavruyan E., Hayrapetyan V., Kopaliani N, Tsingarska E., Krofel M, Skoglund P. & Bogdanowicz W. 2019. Global Phylogeographic and Admixture Patterns in Grey Wolves and Genetic Legacy of An Ancient Siberian Lineage. Scientific Reports. (2019) 9: 1732Ninua L, Tarkhnishvili D, Gvazava E. Phylogeography and taxonomic status of trout and salmon from the Ponto-Caspian drainage, with the inference on the European Brown Trout evolution and taxonomy. Ecology and Evolution 2018 Mar; 8(5): 2645–2658
5. Divya Venkatesh , Marjolein J. Poen, Theo M. Bestebroer , Rachel D. Scheuer, Oanh Vuong , Mzia Chkhaidze , Anna Machablishvili , Jimsher Mamuchadze , Levan Ninua, Nadia B. Fedorova , Rebecca A. Halpin , Xudong Lin , Amy Ransier , Timothy B Stockwell , David E. Wentworth, Divya Kriti, Jayeeta Dutta , Harm van Bakel , Anita Puranik , Marek J Slomka , Steve Essen , Ian H. Brown, Ron A.M. Fouchier , Nicola S. Lewis. Avian influenza viruses in wild birds: virus evolution in a multi-host ecosystem. Journal of Virology, 16 May 2018
6. Alexander Gavashelishvili, Yuriy A Yarovenko, Elmar A. Babayev, Giorgi Mikeladze, Zurab Gurielidze, Davit Dekanoidze, Niko Kerdikoshvili, Levan Ninua, Nika Paposhvili. Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus. Journal of Mammalogy, xx(x):1–13, 2018 
7. Kopaliani N, Gurielidze Z, Ninua L. 2017.Records of anomalously white harbour porpoises and atypical pigmented short-beaked common dolphin in the Georgian Black Sea waters. Journal of Black Sea/ Mediterranean Environment http://www.blackmeditjournal.org/index.php/component/k2/item/575
8. Shvelidze T (2016). First Record of Oithona davisae Ferrari and Orsi, 1984 (Copepoda, Cyclopoida) and Seasonal Variations in the Georgian Black Sea Coast. Acad. J. Sci. Res. 4(11): 424-428.
9. Rutkowski R, Krofel M, Giannatos G, Ćirović D, Männil P, Volokh A., Lanszki J, Heltai M, Ovidiu C. Banea, Hayrapetyan V, Kopaliani N , Miliou A.,. Tryfonopoulos G., Lymberakis P, Penezić A., Pakeltytė G., Suchecka E., Bogdanowicz W. 2015. A European Concern? Genetic Structure and Expansion of Golden Jackals (Canis aureus) in Europe and the Caucasus. PLOS ONE. November 5, 2015
10. Mallon D., Weinberg P. and N. Kopaliani. Status of the Prey Species of the Leopard in the Caucasus. Cat News Special Issue. No 2. IUCN. Species Survival Commission.Cat Specialis Group. Stampfli Publikationen AG, Bern,Switzerland. 2007.
11. Lukarevsky V., Akkiev M., Askerov E., AgiliA., Can E., Gurielidze Z, Kudaktin A., Malkhasyan A and Yarovenko Y. Status of the Leopard in the Caucasus. Cat News Special Issue. No 2. IUCN. Species Survival Commission.Cat Specialis Group. Stampfli Publikationen AG, Bern, Switzerland. 2007.
12. Kopaliani N., Gurielidze Z. Status of Turs in Georgia and Conservation Action Plan. Status and Protection of Globally threatened species in the Caucasus. CEPF /WWF. 2009.
13. Murtskhvaladze M., Gurielidze Z., Kopaliani N., Tarkhnishvili D. Gene introgression between Gazella subguturrosa and G. marica: limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica. 4, 383-386. 2012.
14. Bogdanowicz W, Dabrowski M, Hayrapetyan V, Kopaliani N, Pilot M, Tsingarska E, Yavruyan E. Genetic variability of the grey wolf Canis lupus in the Caucasus in comparison with Europe and the Middle East: Distinct or intermediary population? PLOS ONE . 2014
15. Kopaliani N, Shakarashvili M, Gurielidze Z, Qurkhuli T and Tarkhnishvili D. Hybridization of wolf and dog in Georgia (Caucasus): introgression of wolf alleles into shepherd dog population . The Journal of Heredity. 2014.
16. ზ. გურიელიძე, ნ. კოპალიანი, ნ. დევიძე, მ. შაქარაშვილი, შავი ზღვის ბიომრავალფეროვნება. სიტუაციის ანალიზი (ქართულად და ინგლისურად). GIZ. 96 გვ. 2012.

მაღალმთის ეკოსისტემების კვლევის პროგრამა 

2006-2020 წწ.

რეფერირებად საერთაშორისო პერიოდულ გამოცემებში

1. Jolokhava T., Abdaladze O., Gadilia Sh., Kikvidze Z. (2020): Variable soil pH can drive changes in slope aspect preference of plants in alpine desert of the Central Great Caucasus (Kazbegi district, Georgia). Acta Oecologica 105, 103582 ISI
2. Kikvidze Z., Jolokhava T., Bakhia A., Abdaladze O. (2020): Jumping the barrier: does a glacier tongue affect species distribution along the elevation gradient in the subnival and nival belts? A case study on Mt. Kazbegi, Georgia, Central Great Caucasus Mountains. Botanica Serbica 44(2): 219-229. 
3. Jolokhava T., Abdaladze O., Asanidze Z., Kikvidze Z. (2020): Distribution of plant species and the rock particle size in subnival habitats of the Central Great Caucasus. Soil & Environment. 39(2): 194-203
4. Buchner, J., Yin, H., Frantz, D., Kuemmerle, T., Askerov, E., Bakuradze, T., Bleyhl, B., Elizbarashvili, N., Komarova, A., Lewińskaa, K.E., Rizayeva, A., Sayadyan, H., Bin Tank, B., Tepanosyan, G., Zazanashvili, N., Radelof, V.C. (2020): Land-cover change in the Caucasus Mountains since 1987 based on the topographic correction of multi-temporal Landsat composites. Remote Sensing of Environment, 248:111967
5. Bussman R. W., Zambrana N. P., Sikharulidze Sh., Kikvidze Z., Darchidze M., Ekhvaia J. et al. (2020): From the sea to the mountains: plant use in Ajara, samegrelo and Kvemo Kartli, Sakartvelo (republic of Georgia), Caucasus. Ethnobotany Research and Applications, 20 (9).
6. Murtazaliev R., Anatov Dzh., Ekhvaia J., Guseinova Z., Batsatsashvili K. (2020): Intraspecific variability of some functional traits of Trigonocaryum involucratum (Steen) Medw., a Caucasus endemic plant. Botanica Serbica, 44(2): 129-136. 
7. Mosulishvili M., Bedoshvili D., Maisaia I., Chkhutiashvili G. & N. Ustiashvili (2020): Aegilops tauschii, the D-genome donor of Triticum and the geographic origin of hexaploid wheat. Annals of Agrarian Science, 18, 4
8. Bedoshvili D., Mosulishvili M., Chkhutiashvili G., Chokheli M., Maisaia I. & N. Ustiashvili (2020):  Heritage wheats of Georgia. Annals of Agrarian Science, 18, 2: 123 - 129
9. Mosulishvili M., Bedoshvili D., Maisaia I., Rusishvili N., Chkhutiashvili G. & M. Merabishvili (2020): Diversity of wheat in modern Georgia versus discoveries of the archaeological excavations of the 6^th millennium BC Neolithic sites of Lower Kartli region. Annals of Agrarian Science 19, 1 
10. Kazancı C., Oruç S., & M. Mosulishvili (2020): Medicinal ethnobotany of wild plants: A cross-cultural comparison around Georgia-Turkey border, The Western Lesser Caucasus. Journal of Ethnobiology and Ethnomedicine
11. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K. and Hart, R.E. (2020): An ethnobotany of Kakheti and Kvemo Kartli, Sakartvelo (Republic of Georgia), Caucasus. Ethnobotany Research and Applications, 19: 1-27. 
12. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Darchidze, M., Manvelidze, Z., Ekhvaia, J., Kikodze, D., Khutsishvili, M., Batsatsashvili, K. and Hart, R.E. (2020): From the sea to the mountains-plant use in Adjara, Samegrelo and Kvemo Svaneti, Sakartvelo (Republic of Georgia), Caucasus. Ethnobotany Research and Applications, 20: 1-34. 
13. Goginashvili N., Ekhvaia J., Dobordjginidze R., Bachilava M., Tvauri I., Kobakhidze N. (2020): Comparative analysis of the three Caucasian oak taxa in Georgia (South caucasus) based on leaf morphological variation. Annals of Agrarian Sciences, 18(8)
14. Kuemmerle, T., Bluhm, H., Ghoddousi, A., Arakelyan, M., Askerov, E., Bleyhl B., Ghasabyan, M., Gavashelishvili A., Heidelberg A., Malkhasyan A., Manvelyan K., Soofi M., Yarovenko Yu., Weinberg P. & Zazanashvili N. (2020): Identifying priority areas for restoring mountain ungulates in the Caucasus ecoregion. Conservation Science and Practice. 276
15. Silakadze N., Kilian N., Korotkova N., Mosulishvili M. & T. Borsch. (2019): Multiple evolution of high mountain bellflowers with solitary flowers and calyx scales only renders a core of Campanula subg. Scapiflorae as a Caucasian clade. Systematics and Biodiversity, Volume 16, 4: 2-67 
16. Pistrick K., Akhalkatsi M., Ekhvaia J. (2019): Plant genetic resources of higher altitudes in southwestern Georgia (Samtskhe-Javakheti and Ajara). Genetics Resouces and Crop Evolution. 
17. Bleyhl B., Arakelyan M., Askerov E., Bluhm H., Gavashelishvili A., Ghoddousi A., Heidelberg A., Khorozyan I., Malkhasyan A., Manvelyan K., Masoud M., Moqanaki E., Radeloff V.C., Soofi M., Weinberg P., Zazanashvili N. and Kümmerle T. (2019): Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Diversity and Distributions 25: 129-141
18. Askerov, E., Talibov, T., Manvelyan, K., Zazanashvili, N., Fatullayev, P. & Malkhasyan, A. (2019): Leopard (Panthera pardus) reoccupying its historic range in the South Caucasus: a first evidence (Mammalia: Felidae). Zoology in the Middle East 1-2: 88-90
19. Mosulishvili, M., D. Bedoshvili, I. Maisaia, G. Chkhutiashvili (2019): Georgia, the South Caucasus as the homeland of the hexaploid wheat. Annals of Agrarian Science. 17, 3: 287- 297
20. Zenderland J., Hart R., Bussmann R. W., Paniagua Zambrana N. Y., Sikharulidze Sh., Kikvidze Z., Kikodze D., Tchelidze D., Khutsishvili M., Batsatsashvili K. (2019): The Use of “Use Value”: Quantifying Importance in Ethnobotany. Economic Botany. 1-11

21. Kazanci C., Oruç S., Mosulishvili M. (2019): Cultural Keystone Species Without Boundaries: A Case Study on Wild Woody Plants of Transhumant People Around the Georgia-Turkey Border (the Western Lesser Caucasus). Journal of Ethnobiology. 39, 4: 1-30 

22. Ekhvaia J., Simeone M. C., Silakadze N., Abdaladze O. (2018): Morphological diversity and phylogeography of the Georgian durmast oak (Q. petraea subsp. iberica) and related Caucasian oak species in Georgia (South Caucasus). Tree Genetics & Genomes 14 (2), 17
23. Martín-Benito D., Pederson N., Köse N., Doğan M., Bugmann H., Mosulishvili M. & Ch. Bigler. (2018): Pervasive effects of drought on tree growth across a wide climatic gradient in the temperate forests of the Caucasus. Global Ecology and Biogeography 27: 1314-1325
24. Bussmann R. W., Paniagua Zambrana N. Y.,Sikharulidze Sh., Kikvidze Z., Kikodze D., Tchelidze D., Batsatsashvili K., Hart R. E., Robbie E. (2018): Unequal brothers-Plant and fungal use in Guria and Racha, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 17, no. 1 (2018): 7-33
25. Kupradze I., Inashvili T., Batsatsashvili K., Lachashvili N., Gabelashvili S. (2018): Lichens of the arid region of David Gareji, Georgia (South Caucasus). Herzogia 31, no. 1: 268-275
26. Brander, L.M., Tankha, S. Sovann, G., Sanadiradze, G., Zazanashvili, N. Kharazishvili, D., Memiadze, N., Osepashvili, I., Beruchashvili, G., Arobelidze, N. (2018): Mapping the economic value of landslide regulation by forests. Ecosystem Services, vol. 32, part A., August 2018, pp. 101-109. 
27. Bleyhl, B., Arakelyan, M., Askerov, E., Bluhm, H., Gavashelishvili, A., Ghasabian, M., Ghoddousi, A., Hedelberg, A., Khorozyan, I., Malkhasyan, A., Manvelyan, K., Masoud, M., Moqanaki, E.M., Radeloff, V.C., Soofi, M., Weinberg, P., Zazanashvili, N. and Kuemmerle, T. (2018): Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Biodiversity and Distribution. First published: 25 September 2018 
28. Nakhutsrishvili G. & Abdaladze O. (2017): Vegetation of the Central Great Caucasus along W-E and N-S Transects. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 11-16.
29. Nakhutsrishvili G. & Abdaladze O. (2017): Plant Diversity of the Central Great Caucasus. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 17-112.
30. Nakhutsrishvili, G. & Batsatsashvili, K. (2017): Quantitative analysis of the phytosociological relevés from the Central Greater Caucasus. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 113-132.
31. Randin Ch. F., Pellissier L., Guisan A., Nakhutsrishvili G. (2017): A Comparison of Climatic Niches of the same Alpine Plant Species in the Central Caucasus and the Alps. . In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 133-144.
32. Nakhutsrishvili, G., Batsatsashvili, K., Rudmann-Maurer, K., Körner, Ch. & Spehn, E. (2017): New Indicator values for Central Caucasus flora. In: Nakhutsrishvili, G. Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment, Springer, pp. 145-160.
33. Bussmann, R.W., Batsatsashvili, K., Kikvidze, Z., Khutsishvili, M., Maisaia, I., Sikharulidze, Sh.,Tchelidze, D., Fayvush, G., Alexanyan, A., Mehdiyeva, N.P., Alizade, V.M., Paniagua Zambrana, N. Y. 2017. Ethnobotany of the Caucasus. Bussmann, R.W. (ed.). Springer. 708 p. 
34. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K., Hart, R.E. and Pieroni, A. (2017): Your poison in my pie - the use of Potato (Solanum tuberosum L.) leaves in Sakartvelo, Republic of Georgia, Caucasus, and Gollobordo, Eastern Albania. Economic Botany 
35. Mosulishvili M., Maisaia I., Batsatsashvili K. (2017): Proposal to conserve the name Triticum palaeocolchicum against T. karamyschevii (Poaceae). Taxon, 66, 12: 519-521 
36. Mumladze, L., Asanidze, Z., Walther, F. and Hausdorf, B. (2017): Beyond elevation: testing the climatic variability hypothesis vs. Rapoport’s rule in vascular plant and snail species in the Caucasus. Biological Journal of the Linnean Society, p.blx027 
37. Mumladze, L., Ulrich, W., Asanidze, Z. and Japoshvili, G. (2017): An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Écoscience, pp.1-5
38. Mosulishvili M., Bedoschvili D., I. Maisaia. (2017): A consolidated list of Triticum species and varieties of Georgia to promote the repatriation of local diversity from foreign genebanks. Annals of Agrarian Science. 15, 1: 61-70.
39. Bakuradze T., Kandaurov A., Mosulishvili M., Nikolaishvili D., Gvilava M. & S. Kenkebashvili. (2017): Spatial Analysis Used in Baseline Study for the Preparation of Management Plan of Kazbegi Protected Areas. Earth Sciences. Special Issue: New Challenge for Geography: Landscape Dimensions of Sustainable Development. Vol. 6, No 5-1, pp. 93-110 
40. Bleyhl, B., Baumann, M., Griffiths, P., Heidelberg, A., Manvelyan, K., Radeloff, V., Zazanashvili, N., Kuemmerle, T. (2017): Assessing landscape connectivity for large mammals in the Caucasus using Landsat 8 seasonal image composites. Remote Sensing of Environment, vol. 193:193-203
41. Breitenmoser, U., Askerov, E., Soofi, M., Breitemoser-Wursten, Heidelberg, A., Manvelian, K., and Zazanashvili, N. (2017): Transboundary leopard conservation in the Lesser Caucasus and Alborz Range. Cat News 65, pp. 24-25     http://fungi.biodiversity-georgia.net/ 
42. Winkler M., A. Lamprech, K. Steinbauer, K. Hülber, J-P. Treurillat, F. Breiner, P. Choler, S. Ertl, G.A. Giron, G. Rossi, P. Vittoz, M. Akhalkatsi, Ch. Bay, J.L. Alonso, T. Bergström, M.L. Carranza, E. Corcket, J. Dick, B. Ershbamer, R. F. Calzado, A. M. Forsaa, R. G. Gavilan, D. Chosn, Kh. Gigauri, D. Huber, R. Kanka, G. Kazakiz, M. Klipp. J. Kollar, T. Kudernatsch, P. Lersson, M. Mallaun, O. Michelsen, P. Moiseev, D. Moiseev, Ulf, Mollau, J.M. Mesa, U.M. di Cella, L. Nagy, M. Petey, M. Puscas, Ch. Rixen, A. Stanisci, M. Suen, A. O. Syverhuset, M. Tomaselli, P. Unterluggauer, T. Ursu, L. Villar, M. Gottfried and H. Pauli (2016): The rich sides of mountain summits-a pan-European view on aspect preferences of alpine plants. Journal of Biogeography 43: 2261-2273. 
43. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. (2016): Alpine plant distribution and thermic vegetation indicator on GLORIA summits in the Central Greater Caucasus. Pakistan Journal of Botany, 48(5): 1893-1902 
44. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K. & Hart, R.E. (2016): A comparative ethnobotany of Khevsureti, Samtskhe-Javakheti, Tusheti, Svaneti, and Racha-Lechkhumi, Republic of Georgia (Sakartvelo), Caucasus. Journal of Ethnobiology and Ethnomedicine, 12(1), p.43.
45. The impacts of management and site conditions on the phytodiversity of the upper montane and subalpine belts in the Central Greater Caucasus. Tephnadze N., Abdaladze O., Nakhutsrishvili G., Simmering D., Waldhardt R., Otte A. Phytocoenologia, vol. 44, #3-4, pp. 255-291(37). 
46. Genetic relationships between wild progenitor pear (Pyrus L.) species and local cultivars native to Georgia, South Caucasus. Asanidze, Z., Akhalkatsi, M., Henk, A.D., Richards Ch. M., Volk G. M. Flora, 209, pp. 504-512. 2014
47. Tephnadze N., Abdaladze O., Nakhutsrishvili G., Simmering D., Waldhardt R., Otte A. (2013): Estimating the relative importance of land management and abiotic site conditions for phytodiversity in the Kazbegi district, Georgia. - Phytocoenosis
48. Cavieres, L., Pugnaire, F., Schöb, C., Xiao, S., Anthelme, F., Björk, R., Dickinson, K., Cranston, B., Gavilán, R., Gutiérrez-Girón, A., Kanka, R., Kikvidze Z., Maalouf, J.-P., Mark, A., Noroozi, J., Parajuli, R., Phoenix, G., Reid, A., Ridenour, W., Rixen, C., Wipf, S., Zhao, L., Escudero, A., Zaitchik, B., Aschehoug, E., Lingua, E. and Callaway, R. (2013): Facilitative plant interactions and climate simultaneously drive alpine plant diversity. – Ecology Letters 
49. Jimbo, K., Kubota, K. & Kikvidze, Z. (2013): Ground beetle succession on Mount Fujisan. – Biogeography (Japan). 15
50. Asensio, C. Lozano, F.J., Ortega, E. & Kikvidze, Z. (2013): Study on the effectiveness of an agricultural technique based on aeolian deposition, in a semiarid environment. Environmental Engineering and Management Journal 12
51. Ekhvaia J., Gurushidze M, Blattner F.R., Akhalkatsi M. (2013): Genetic diversity of Vitis vinifera in Georgia: relationships between local cultivars and wild grapevine, V. vinifera subsp. sylvestris. - Theoretical and applied Genetics 
52. Kürschner, H., Batsatsashvili, K. & Parolly, G. 2013. Noteworthy additions to the bryophyte flora of Georgia. – Herzogia 26: 213 –216.
53. Montalvo Mancheno, C.S., Nakhutsrishvili, G. & Zazanashvili, N. (2013): A Perspective on Ecological Corridor for Maintaining Healthy Ecological Processes in the Caucasus. Research in Protected Areas, Mittersill, pp. 523-526.
54. Barjadze, Sh., Asanidze, Z. (2013): Morphometric investigation of grapevine and willow root-feeding aphid, Pemphigus saliciradicis (Börner, 1950) (Hemiptera: Aphididae: Eriosomatinae). - Dtsch. Entomol. Z. 60 (1), 45–52 
55. Butterfield, B.J., L. A. Cavieres, R. M. Callaway, B. J. Cook, Z. Kikvidze, C. J. Lortie, R. Michalet, F. I. Pugnaire, C. Sch€ob, S. Xiao, B. Zaitchek, F. Anthelme, R. G. Bj€ork, K. Dickinson, R. Gavilan, R. Kanka, J.-P. Maalouf, J. Noroozi, R. Parajuli, G. K. Phoenix, A. Reid, W. Ridenour, C. Rixen, S. Wipf, L. Zhao, R. W. Brooker (2013): Alpine cushion plants inhibit the loss of phylogenetic diversity in severe environments. - Ecology Letters, 1-8. 
56. Miyata M., Wu B., Kikvidze Z., Fukuda K. (2013): Mycorrhizal Cirsium purpuratum improves the growth of Clematis stans in volcanic scoria of Mount Fuji. - Micoscience, vol. 54, (4): 279-284.
57. Khojasteh F., M. A. Z. Chahouki, H. Azarnivand, Z. Kikvidze (2013): Life form and preference can drive spatial relationships among plant species in semi-arid rangelands of middle Iran. - The Rangeland Journal. 
58. Suzuki M, Miyashita T, Kabaya H, Ochiai K, Asada M, & Kikvidze Z. (2012): Deer herbivory as an important driver of divergence of ground vegetation communities in temperate forests. - Oikos. 
59.  Abukava K, Yamamuro M, Kikvidze Z, Asada A, Xu C, & Sugimoto K. (2012): Assessing the biomass and distribution of submerged aquatic vegetation using multibeam echo sounding in Lake Towada, Japan. - Limnology
60. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. (2013): Monitoring of vascular plant diversity in the alpine zone of the Central Greater Caucasus in a changing climate Turkish Journal of Botany, 37: 1104-1114. 
61. Pauli H., Gottfried M., Dullinger S., Abdaladze O., Akhalkatsi M., Benito Alonso J. L., Coldea G., Dick J., Erschbamer B., Calzado R. F., Ghosn D., Holten J. I., Kanka R., Kazakis G., Kollár J., Larsson P., Moiseev P., Moiseev D., Molau U., Mesa J. M., Nagy L., Pelino G., Puşcaş M., Rossi G., Stanisci A., Syverhuset A. O., Theurillat J-P., Tomaselli M., Unterluggauer P., Villar L., Vittoz P., Grabherr G. (2012): Recent plant diversity changes on Europe’s mountain summits. Science vol. 336, pp. 353-355. 
62. Gottfried, M., Pauli, H., Futschik, A., Akhalkatsi, M., Barančok P., Alonso, J.L.B., Coldea, G., Dick, J., Erschbamer, B., Rosa, M., Calzado, F., Kazakis G., Krajči, J., Larsson, P., Mallaun, M., Michelsen, O., Moiseev, D.15, Moiseev, P., Molau, U., Merzouki, A., Nagy, L., Nakhutsrishvili, G., Pedersen, B., Pelino, G., Puscas, M., Rossi, G., Stanisci, A., Theurillat, J.-P., Tomaselli, M., Villar, L., Vittoz, P., Vogiatzakis, I., Grabherr G. (2012): An early warning signal for a continental-scale response to ongoing climate change. Nature Climate Change 2, 111–115.
63. Caprez, R., Spehn, E., Nakhutsrishvili G., Körner Ch. (2011): Drought at erosion edges selects for a ‘hidden’ keystone species. Plant Ecology & Diversity. ISSN: 1755-0874. 
64. Asanidze, Z., Akhalkatsi, M., Gvritishvili, M. (2011): Comparative morphometric study and relationships between the Caucasian species of wild pear (Pyrus spp.) and local cultivars in Georgia. Flora 206, 11
65. Nakhutsrishvili G. (2011): La vegetation des zones de haut montagne du Caucase. Ethnopharmacologia. Nr. 47: 45-51. 
66. Nakhutsrishvili G. (2011): Historique de Institute de Botanique de Georgie. Ethnopharmacologia. Nr. 47: 39-44. 
67. Akhalkatsi M., Ekhvaia, J., Mosulishvili M., Nakhutsrishvili G., Abdaladze O., Batsatsashvili K. (2010): Reasons and processes leading to the erosion of crop genetic diversity in mountainous regions of Georgia. Mountain Research and Development. 30, 3:304-310. 
68. Ekhvaia J., Akhalkatsi M. (2010): Morphological variation and relationships of Georgian populations of Vitis vinifera L. subsp. sylvestris (C.C. Gmel.) Hegi. Flora 205, 9:608-617.
69. Inashvili, Tz. & Batsatsashvili, K. (2010): New Lichen Records from Georgia. Turkish Journal of Botany. 34(6): 549-553. 
70. Erschbammer B., Mallaun M., Unterluggauer P., Abdaladze O., Akhalkatsi M., Nakhutsrishvili G. (2010): Species richness and distribution above the treeline in the Central Alps (Italy) and in the Central Caucasus (Georgia). Tuexenia. 30:11-29. (ISSN: 0722-494X) 
71. Hughes N.M., Johnson D.M., Akhalkatsi M., Abdaladze O. (2009): Characterizing Betula litwinowii Seedling Microsites at the Alpine-Treeline Ecotone, Central Greater Caucasus Mountains, Georgia. – Arctic, Antarctic & Alpine Research, vol. 41, No 1, pp. 112-118. 
72. Dullinger S., I. Kleinbauer, H. Pauli, M. Gottfried, Ch. Klettner, R. Brooker, L. Nagy, J.-P. Theurillat, J. I. Holten, O. Abdaladze, J-L. Benito, J.-L. Borel, G. Coldea, D. Ghosn, R. Kanka, A. Merzouki, P. Moiseev, U. Molau, K. Reiter, G. Rossi, A. Stancisi, M. Tomaselli, P. Unterlugauer, and G. Grabherr (2007): Do co-occurrence patterns in alpine plant communities indicate a stress-driven increase in facilitation? - Journal of Ecology.  
73. Akhalkatsi M., Abdaladze O., Nakhutsrishvili G., Smith W. K. (2006): Rhododendron caucasicum and Microtopography Extend the Betula litwinowii Alpine Treeline (Caucasus Mountains, Georgia). – Arctic, Antarctic & Alpine Research, vol. 38, #4, pp. 441-448. ISSN: 1523-0430

სხვა პუბლიკაციები 

1. Zazanashvili N., M. Garforth and G. Nakhutsrishvili (2021): Caucasus-Anatolian-Hyrcanian Temperate Forests. Chapter 50035 for the Encyclopedia of Conservation.  Elsevier
2. Jolokhava T., Kikvidze Z., Gigauri Kh., Abdaladze O. (2021): Maximum soil temperature, plant available NPK, pH and soil organic matter (SOM) along the altitudinal gradient (3000-3900 m a.s.l.) in the Central Great Caucasus. 
3. Burgaz A.R., Teuvo A., Inashvili T., Batsatsashvili K., Kupradze I. (2018): Study of Georgian Cladoniaceae. Botanica complutensis 42: 19-55
4. Maharramova, E., Moqanaki, E.M., Askerov, E., Faezi, S., Alinezhad, H., Mousavi, M., Kuemmerle, T., Heidelberg, A. & Zazanashvili, N. (2018): Transboundary leopard movement between Azerbaijan and Iran in the Southern Caucasus. Cat News 67: 8-10
5. Kuemmerle, T., Blehyl, B., Bluhm, H., Ghoddousi, A., Heidelberg, A., Sarukhanova, S., Zazanashvili, N. (2018): Identifying priority areas for restoring large mammal populations in the Caucasus Ecoregion. 5th European Congress of Conservation Biology, 12-15 June, Jyväscylä, Finland 
6. Gavrilović M., Tešević V., Đorđević I., Rajčević N., Bakhia A., Garcia Jacas N., Susanna A., Marin P. D., Janaćković P. (2018): Leaf micromorphology, antioxidative activity and a new record of 3-deoxyamphoricarpolide of relict and limestone endemic Amphoricarpos elegans Albov (Compositae) from Georgia. Archives of Biological Sciences 2018 OnLine-First Issue 00, Pages: 22-22
7. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2017): Ethnobotany of Samtskhe-Javakheti, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 16 (1):7-24 
8. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2017): Plants in the spa – the medicinal plant market of Borjomi, Sakartvelo (Republic of Georgia), Caucasus. Indian journal of traditional knowledge 16 (1): 25-34
9. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze Sh., Kikvidze Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K., Hart, R.E. and Robbie, E. (2018): Unequal brothers–Plant and fungal use in Guria and Racha, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 17(1): 7-33
10. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2016): Medicinal and Food Plants of Svaneti and Lechkhumi, Sakartvelo (Republic of Georgia), Caucasus. Medicinal and Aromatic Plants (Los Angel), 5: 266 
11. Abdaladze O., Nakhutsrishvili G., Batsatsashvili K., Gigauri Kh., Jolokhava T., Mikeladze G. (2015): Sensitive Alpine Plant Communities to the Global Environmental Changes (Kazbegi Region, the Central Great Caucasus). - American Journal of Environmental Protection. Special Issue. Applied Ecology: Problems, Innovations. Vol. 4, No.3-1, pp. 93-100.
12. Askerov E., Talibov T., Manvelyan K., Zazanashvili N., Malkhasyan A., Fatullayev P. & Heidelberg A. (2015): South-Eastern Lesser Caucasus: the most important landscape for conserving the Leopard (Panthera pardus) in the Caucasus region (Mammalia: Felidae). - Zoology in the Middle East,v. 61, 2.
13. Kupradze I., Jorjadze I., A. Arabidze, T. Beltadze, Batsatsashvili K. (2015): Ethnobiological Study of Svaneti Fungi and Lichens: History of Research, Diversity, Local Names and Traditional Use. – American Journ. of Environmental Protection. Special Issue. Applied Ecology: Problems, Innovations. Vol. 4, No. 3-1, pp. 101-110.
14. Akhalkatsi M., Asanidze Z. (2015): Pyrus caucasica Fed. Ancestor of Georgian Pear Landraces. – Science, technology and development, 34 92), pp. 105-108.
15. Wine, Beer, Snuff, Medicine, and Loss of Diversity-Ethnobotanical travels in the Georgian Caucasus. Bussmann, R. W., Paniagua Zambrana N. Y., Sikharulidze Sh., Kikvidze Z., Kikodze D., Jinjikhadze T., Shanshiashvili T., Chelidze D., Batsatsashvili K. & Bakanidze N. Ethnobotany Research and Applications, 12: 237-313, 2014
16. Caucasus endemic vascular plant species Red List assessments (385 taxa). Scharz, G.E., Shulkina, T.V., Nakhutsrishvili, G., Batsatsashvili, K. et al. (evaluators). The IUCN Red List of Threatened Species, 2014
17. Vascular plant diversity and climate change in the alpine zone of the Central Greater Caucasus. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. International Journal of Ecosystems and Ecology Science. ISSN 2224-4980, v. 4 (4), pp. 573-589, 2014
18. Waldhart R., Abdaladze O., Otte A., Simmering D. (2011): Ursachen und Folgen des Landschaftswandel im Kaukasus Georgiens: Interdisziplinäre Forscung Für eine nachhaltigere Zukunft. Spiegel der Forschung. 28, Nr 2: 4-15
19. Otte A., Akhaltkatsi, M., Nakhutsrishvili, G., Simmering, D., Waldhardt, R. (2011): Phytodiversität in Georgien: Die Bedeutung von Standort und Landnutzung im Großen und Kleinen Kaukasus. Spiegel der Forschung 28, Nr 2:24-31
20. Pistrick K., Akhalkatsi M., Girgvliani T., Shanshiashvili T. (2010): Sammlung von Saat- und Pflanzgut pflanzengenetischer Ressourcen in Svanetien und Lečchumi (Georgien, Kaukasus) 2009. Berichte Ges. Pflanzenbauwiss. 5, 1:125-127
21. Mosulishvili M., Arabuli G. (2010): Buxus Stands in East Georgia. Bull. of Georg. Nation. Museum, #2, pp. 128-132. 
22. Lachashvili N., Khetsuriani L., Batsatsashvili K. (2010): Effect of grazing on some communities of Bothriochloeta ischaemum in Shida Kartli (East Georgia). Bull. of Georgian National Academy of Sciences. 8, #1-2, pp. 20-25.
23. Nakhutsrishvili G., Akhalkati M., Abdaladze O. (2009): Main Treats to the Mountain Biodiversity in Georgia (the Caucasus). – Mountain Forum Bulletin, vol. IX, Issue 2, pp. 18-19.
24. Pistrick, K., Akhalkatsi M., Girgvliani T. Shanshiashvili T. (2009): Collecting plant genetic resources in Upper Svaneti (Georgia, Caucasus mountains). J. Agric. Rural Dev. Trop. Subtrop. Supplement 92:127-135.
25. Schultzke, R., Zazanashvili, N. and Osepashvili, I. (2009): Restoration of forest landscapes in the Southern Caucasus. Pp. 119-125 in: Bodegom, V, Jan, A., Savenije, H. and Wit, M. (Eds.). Forest and Climate Change: adaptation and mitigation. Tropendos International, Wegeningen, The Netherlands.
26. Batsatsashvili K., Chelidze N. (2009): Lichens as Bioindicators of Tbilisi Air pollution. TSU Publ., pp. 295–309 (Georgian). 
27. Bohn U., Zazanashvili N., Nakhutsrishvili G. (2007): The Map of the Natural Vegetation of Europe and its application in the Caucasus Ecoregion. – Bulletin of the Georgian National Academy of Sciences, col. 175, #1, pp. 112-121.

სამეცნიერო კრებულებში:

1. Zazanashvili, N., Sanadiradze, G., Garforth, M., Bitsadze, M., Manvelyan, K., Askerov, E., Mousavi, M., Krever, V., Shmunk, V., Kalem, S. and Devranoğlu Tavsel, S., eds. (2020): Ecoregional Conservation Plan for the Caucasus: 2020 Edition. WWF, KfW, Tbilisi.
2. Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. (2020): Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
3. Zazanashvili, N., Manvelyan, K., Askerov, E., Mousavi, M., Krever, V., Kalem, S. & Garforth, M. (2020): The boundaries and bio-physical features of the Caucasus Ecoregion. Pp. 9-20 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
4. Bitsadze, M., Manvelyan, K., Askerov, E., Mousavi, M., Shmunk, V., Kalem, S., Zazanashvili, N., Mnatsekanov, R., Devranoglu Tavsel, S., Cevik, M., Ahmadova, K., Beruchashvili, G., Kandaryan, A. & Garforth, M. (2020): Key Biodiversity Areas in the Caucasus Ecoregion. Pp. 21- 28 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
5. Zazanashvili, N., Manvelyan, K., Askerov, E., Mousavi, M., Shmunk, V., Kalem, S., Beruchashvili, G., Bitsadze, M. & Garforth, M. (2020): Conservation landscapes in the Caucasus Ecoregion. Pp. 29 - 36 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
6. Askerov, E., Manvelyan, K., Gurielidze, Z., Mousavi, M., Shmunk, V., Trepet, S., Kütükçü, A. E., Heidelberg, A. & Zazanashvili, N. (2020): Status of large carnivores in the Caucasus. Pp. 37 - 47 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
7. Weinberg, P., Askerov, E., Manvelyan, K., Gurielidze, Z., Mousavi, M., Kütükçü, A.E., Heidelberg, A. & Zazanashvili, N. (2020): Status of large herbivores in the Caucasus. Pp. 48 - 71 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
8. Abdaladze O. (2011): Functional Diversity of Alpine Plants of Georgia and Climate Global Change. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Science of Georgia, Tbilisi: 15-18. 
9. Nakhutsrishvili G. (2011): Actual Problems of Biodiversity. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Scien. of Georgia, Tbilisi: 133-134. 
10. Gigauri Kh., Akhalkatsi M., Nakhutsrishvili G., Abdaladze O. (2011): First Signs of Weak Thermophilisation on the GLORIA-EUROPE Target Region in the Central Caucasus. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Scien. of Georgia, Tbilisi: 19-22.
11. Sharikadze Kh., Nakhutsrishvili G., Akhalkatsi M., Abdaladze O. (2011): Species Diversity of Eroded Slopes in the Kazbegi region. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Scien. of Georgia, Tbilisi: 29-32. (Georgian with English summary).
12. Zazanashvili N., Gavashelishvili A., Montalvo K., Beruchashvili G. (2011): Modelling the Response of South Caucasus Forest Classes to Climate Change. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Scien. of Georgia, Tbilisi: 210-212.
13. Akhalkatsi M., Fritsch R. M., Nakhutsrishvili G., Pistrik K. (2008): Habitats of Allium species in Georgia.– In: M. Keusgen & M. Fritsch (eds.): Botany, Taxonomy and Phytochemistry of Wild Allium L. Species of the Caucasus and Central Asia. Marburg-Gatersleben, pp. 147-156
14. Pistrik K., Akhalkatsi M., Nakhutsrishvili G. (2008): Collecting wild Allium species in Georgia. – In: M. Keusgen & M. Fritsch (eds.): Botany, Taxonomy and Phytochemistry of Wild Allium L. Species of the Caucasus and Central Asia. Marburg-Gatersleben, pp. 45-52

წიგნები და თავები წიგნებში:

1. Bussmann, R.W., Batsatsashvili, K.,  Kikvidze, Z.,  Paniagua Zambrana, N.Y., Khutsishvili, M., Maisaia, I., Sikharulidze, Sh.,Tchelidze, D. et al. (2020): Ethnobotany of the Mountain Regions of Far Eastern Europe. Batsatsashvili, K., Kikvidze, Z. Bussmann, R.W. (eds.). Springer. 1063p.
2. Bussmann, R.W., Batsatsashvili, K.,  Kikvidze, Z. et al. (2020): Ethnobotany of the Mountain Regions of Central Asia and Altai. Batsatsashvili, K., Kikvidze, Z. Bussmann, R.W. (eds.). Springer. 881p.
3. Mosulishvili M., Bedoshvili D., Maisaia I., Rusishvili N., Chkhutiashvili G. (2019): The Ancient, Traditional Georgian Wheat Species and Their Role in Wheat Evolution. in: Proceedings of the International Conference “Wheat in European countries and Georgia as one of the center of origin of Domesticated Wheat”: 232 – 239. Tbilisi, Georgia.
4. Gambashidze G., Jolokhava T. et al. (2019): Soil erosion modelling using RUSLE 1 in Shida Kartli Region, Georgia. Food and Agriculture Organization of the United Nations, Rome, Italy.
5. ნახუცრიშვილი გ., აბდალაძე ო. (2018): ბოტანიკური და ეკოლოგიური კვლევები ყაზბეგის რეგიონში. ნ. ანთიძე (რედ.) მუზეუმი და კულტურული მემკვიდრეობა. თბილისი, გვ. 205–210.
6. გოგინაშვილი, ნ., თვაური, ი., მანველიძე, ზ., მემიაძე, ნ., ასანიძე, ზ., ღამბაშიძე, გ. (2018): აჭარის ტყის იშვიათი მერქნიანი სახეობების ინვენტარიზაციის მასალები. საქართველოს გარემოსა და სოფლის მეურნეობის სამინისტროს სოფლის მეურნეობის სამეცნიერო კვლევითი ცენტრი, გამომც. „უნივერსალი“, თბილისი.
7. Nakhutsishvili G., Batsatsashvili K., Gagnidze R., Shetekauri Sh., Manvelidze Z., Memiadze N. & D. Kharazishvili (2013): Georgia. – In: Schatz, G., Shulkina, T. & Solomon, J. (eds.) Red List of the Endemic Plants of the Caucasus Region. Missouri Botanical Garden Press, Saint Louis, USA 
8. Nakhutsrishvili, G. (2013): The vegetation of Georgia (Caucasus). - Springer-Verlag, Heidelberg New York Dortrecht London, 235 pp.
9. ნ. ელიზბარაშვილი, ნ.ზაზანაშვილი, დ. ნიკოლაიშვილი, ლ. მაჭავარიანი, გ. მელაძე, გ. დვალიშვილი. საქართველოს ბუნება. გამომც. „პალიტრა L.“, თბილისი, 294 გვ. 2013.
10. Nakhutsrishvili, G., Zazanashvili, N. & Batsatsashvili, K. (2011): Why the Colchis and Hyrcan relict forests of the western Eurasian Caucasus should be considered “temperate rainforest”. – In: DellaSala D. (ed.) Temperate and Boreal Rainforests of the World: Ecology and Conservation. Island Press., pp. 214-221.
11. Bondirev, I.V., Seperteladze, Z. KH., Zazanashvili, N.I., Tavartkiladze, A.M., Budagov, B.A., Museibov, M.A., Vardanian, T.G., Balian, A.S. (2009): Critical changes of the ecosystems in Southern Caucasus. Pp. 7-14 in I. V. Bondirev, A. M. Tavartkiladze (Eds.). Critical ecoregions: modern conditions, problems and ways of their decisions. Publishing House. Tbilisi: "Polygraph".
12. Nakhutsrishvili G., Abdaladze O., Akhalkatsi M. (2006): Biotope Types of the Treeline in the Central Greater Caucasus. – In: In: D. Gafta & J. Akereoyd (eds.): “Nature Conservation: Concepts and Practice”, Springer Verlag, pp. 211-225. ISBN 9783540472292.
13. Körner, Ch., Nakhutsrishvili, G., Spehn, E. M. (2006): High-Elevation Land Use, Biodiversity and Ecosystem Functioning. In: Spehn, E. M., Liberman, M., Kőrner, Ch. (Eds.), Land Use Change and Mountain Biodiversity, Taylor & Francis Group, Boca Raton, London, New York, pp. 3-21.
14. Nakhutsrishvili G., Sikharulidze Sh., Abdaladze O. (2006): Bakuriani – Natural and Cultural Resources of the Borjomi Region. – Institute of Botany, Tbilisi, 56 p.