რაოდენობრივი ეკოლოგიის პროგრამა 

2021-2025წწ.

• Tarkhnishvili, D., Todua, M., Iankoshvili, G., Çetintaş, O., Murtskhvaladze, M., & Yanchukov, A. (2025). Collapsing landmasses and secondary contacts: introgression and the evolution of adaptive diversity in Caucasian rock lizards. Evolution, 79(10), 2193-2207.
• Barateli, N., Iankoshvili, G., Tsikolia, N., Seropian, A., & Tarkhnishvili, D. (2025). The tail of the two: a study of twin embryos in Darevskia lizards. Amphibia-Reptilia, 46(2), 241-248.
• Metreveli, V., Kreft, H., Javakhishvili, Z., Akobia, I., Mdivani, S., Chikorashvili, G., & Gavashelishvili, A. (2025). On the relationship between environment and growth of Sweet chestnut (Castanea sativa) in the Caucasus. Frontiers in Forests and Global Change, 8, 1670459.
• Murtskhvaladze, M., Ninua, L., Budagashvili, N., Tevdoradze, E., Gurgenidze, Z., Kotorashvili, A., ... & Javakhishvili, Z. (2025). Tracheal and cloacal bacterial diversity of red-listed Eastern Imperial Eagle (Aquila heliaca). Frontiers in Microbiology, 16, 1477032.
• Kvaratskhelia, R., & Gavashelishvili, A. (2025). Common Yew (Taxus baccata) as a climate archive: Reconstructing 200 years of temperature change in Georgia (Caucasus). Dendrochronologia, 89, 126285.
• Metreveli, V., Kreft, H., Javakhishvili, Z., Akobia, I., Mdivani, S., Chikorashvili, G., & Gavashelishvili, A. (2025). On the relationship between environment and growth of Sweet chestnut (Castanea sativa) in the Caucasus. Frontiers in Forests and Global Change, 8, 1670459.
• CHITADZE, B., DZULIASHVILI, K., & SEROPIAN, A. (2025). New data on harvestmen (Opiliones) of Georgia with the description of a new Rilaena species. Zootaxa, 5725(4), 591-599.
• Seropian, A. (2025). Stuck in between: First case of transverse intersexuality in Trochosa cachetiensis Mcheidze, 1997 (Araneae, Lycosidae) from Georgia. Caucasiana, 4, 153-161.
• Seropian, A., Zamani, A., Bulbulashvili, N., Arsenashvili, E., Zarikian, N., & Szűts, T. (2025). Velvet Revolution: integrative taxonomy of South Caucasian Eresus Walckenaer, 1805 (Araneae, Eresidae), with five new species. Zoological Journal of the Linnean Society, 205(4), zlaf169.
• Seropian, A., Ninua, L., Rostiashvili, I., Bulbulashvili, N., & Makharadze, G. (2025). Corrigendum: New data on spiders (Arthropoda, Arachnida, Araneae) of Georgia with description of a new species from the Tegenaria lyncea group. Caucasiana 4: 65–80. https://doi. org/10.3897/caucasiana. 4. e151922. Caucasiana, 4, 95-96
• Seropian, A., Bulbulashvili, N., Makharadze, G., & Kovács, G. (2025). From burrows to spotlight: first description of the female of Eresus lavrosii Mcheidze, 1997 (Araneae, Eresidae), with notes on the natural history. Caucasiana, 4, 1-14.
• HULSBOSCH, V. P., & SEROPIAN, A. (2025). Burrowed through time: Exploring the enigmatic Cossidae moths of Georgia. Zootaxa, 5701(5), 501-523.Wellenbeck, A., Hein, N., Tarkhnishvili, D., Misof, B., Schmidtlein, S., Janiashvili, Z., ... & Feilhauer, H. (2025). Predicting woody species assemblages using ecophylogenetics and Earth observation data. Forest Ecology and Management, 589, 122763.
• Tielidze, L. G., Mackintosh, A. N., Gavashelishvili, A., Gadrani, L., Nadaraia, A., & Elashvili, M. (2025). Post-Little Ice Age Equilibrium-Line Altitude and Temperature Changes in the Greater Caucasus Based on Small Glaciers. Remote Sensing, 17(9), 1486.
• Kitel, D., & Javakhishvili, Z. (2025). The diet of Barn Owl (Tyto alba) in Georgia. Caucasiana, 4, 105-118.
• Hwang, B. C., Giardina, C. P., Barrios‐Garcia, M. N., Diao, H., Duboscq‐Carra, V. G., Hemp, A., ... & Metcalfe, D. B. (2025). Insect herbivory releases more nutrients in warmer and drier forests.Gl  obal Biogeochemical Cycles, 39(4), e2024GB008367.
• Japaridze, L. G., & Seropian, A. (2025). Unwanted guests: The first records of two established non-native moth species in the Caucasus, with notes on non-native fauna of Georgia. Caucasiana, 4, 137-151
• Rizayeva, A., Nita, M. D., Yin, H., Buchner, J., Kasraee, N., Rogova, N., ... & Radeloff, V. C. (2025). A half-century of land cover changes in the Caucasus derived from Corona spy satellite and Landsat images. Regional Environmental Change, 25(1), 27.
• Lutgen, D., Peona, V., Chase, M. A., Kakhki, N. A., Lammers, F., de Souza, S. G., ... & Burri, R. (2025). A mosaic of modular variation at a single gene underpins convergent plumage coloration. Science, 390(6770), eado8005.
• Farajli, Z., Kittelberger, K. D., Tanner, C. J., Aghababyan, K., Paposhvili, N., Hakiminejad, M., ... & Haraldsson, T. A. (2025). Status and breeding population of the Saker Falcon (Falco cherrug) in the Caucasus Ecoregion: Regional perspectives and conservation challenges. Caucasiana, 4, 25-43.
• Özkoç, Ö. Ü., Paposhvili, N., Aghababyan, K., Petersen, I. K., & Fox, A. D. (2025). Contraction of range, declines in abundance and site-based threats to Velvet Scoter Melanitta fusca breeding in the Caucasus. Wildfowl, 75, 159-184.
• Topaloudis, A., Cumer, T., Lavanchy, E., Ducrest, A. L., Simon, C., Machado, A. P., ... & Goudet, J. (2025). The recombination landscape of the barn owl, from families to populations. Genetics, 229(1), iyae190.
• Mélanie, S., Necmi, A., Claire, A., Dalibor, B., Tatiana, B., Sylvain, B., ... & Jens-Christian, S. (2025). Seeing yew for the forest: a call to action for improving conservation and restoration of the European yew (Taxus baccata L.). Trees, Forests and People, 101093.
• Mebert, K., Igci, N., Karis, M., Yildiz, M., Schmid, P., Gruenewald, J., ... & Kirschey, T. (2025). CRITICALLY ENDANGERED DAREVSKY'S VIPER: INCISIVE DISTRIBUTION DISCOVERIES AND TAXONOmIC CONSIDERATIONS. Herpetological Conservation and Biology, 20(2).
• Mebert, K., Kariş, M., Yildiz, M. Z., Iğci, N., Stümpel, N., Zinenko, O., ... & Elden, J. M. (2025). Threatened steppe vipers of western Asia: Vipera anatolica and Vipera renardi eriwanensis sl—taxonomy, distribution, and conservation. Herpetol Conserv Biol. 2025b (accepted).
• Zamani, A., Seropian, A., Zarikian, N., Bulbulashvili, N., & Szűts, T. (2025). Different, but still the same: integrative taxonomy confirms a new species of Eresus Walckenaer, 1805 (Araneae, Eresidae) from the South Caucasus.ZooKeys, 1249, 1.
• Metreveli, V., Kreft, H., Javakhishvili, Z., Akobia, I., Mdivani, S., Chikorashvili, G., & Gavashelishvili, A. (2025). On the relationship between environment and growth of Sweet chestnut (Castanea sativa) in the Caucasus. Frontiers in Forests and Global Change, 8, 1670459.
• Seropian, A., Ninua, L., Rostiashvili, I., Bulbulashvili, N., & Makharadze, G. (2025). Corrigendum: New data on spiders (Arthropoda, Arachnida, Araneae) of Georgia with description of a new species from the Tegenaria lyncea group. Caucasiana 4: 65–80. https://doi. org/10.3897/caucasiana. 4. e151922. Caucasiana, 4, 95-96
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• Wüster, W., Kaiser, H., Hoogmoed, M. S., Ceríaco, L. M., Dirksen, L., Dufresnes, C., ... & Böhme, W. (2024). How not to describe a species: lessons from a tangle of anacondas (Boidae: Eunectes Wagler, 1830). Zoological Journal of the Linnean Society, 201(4), zlae099.
• Barateli, N., Gabelaia, M., Iankoshvili, G., & Tarkhnishvili, D. (2024). Head shape divergence between parthenogenetic and their paternal bisexual rock lizards in sympatry. Zoologischer Anzeiger.
• Tarkhnishvili, D., Seropian, A., Erhardt, C., Kachlishvili, N., Krammer, H. J., & Hein, N. (2024). How dispersal rates depend on the prey capture strategy: A case study of Georgia's spiders. Ecology and Evolution, 14(5), e11372.
• Tarkhnishvili, D. (2024). Molecular Markers and Taxonomic Explosion in Herpetology: More or Less Robust Taxonomy? True and False Advantages of DNA Markers. International Journal of Zoology and Animal Biology. 7(2): 00569
• Kolev, Z., Novák, J., & Seropian, A. (2024). On the presence of Hipparchia parisatis (Kollar, 1849) in the Republic of Georgia (Lepidoptera, Nymphalidae, Satyrinae). Caucasiana, 3, 263-271.
• Seropian, A. (2024). The first record of Dictis L. Koch, 1872 (Araneae, Scytodidae) in the Caucasus. Caucasiana, 3, 215-221.
• Seropian, A., Bulbulashvili, N., Makharadze, G., & Baznikin, A. (2024). New and interesting spiders (Arachnida, Araneae) from Vashlovani National Park and Chachuna Managed Reserve (Georgia). Caucasiana, 3, 183-213.
• Seropian, A., Bulbulashvili, N., & Krammer, H. J. (2024). Notes on Liocranidae (Arachnida, Araneae, Dionycha) from Georgia with description of a new species. Caucasiana, 3, 171-181.
• Chitadze, B., Bulbulashvili, N., Japaridze, L. G., Drogvalenko, A., Moulet, P., & Seropian, A. (2024). First record of 15 species of Hemiptera (Hexapoda, Insecta) in Georgia. Caucasiana, 3, 127-143.
• Seropian, A., & Mumladze, L. (2024). A new Psammitis species (Araneae, Thomisidae) from an extinct volcano in Georgia with reevaluation of the generic position of Xysticus marmoratus Thorell, 1875. Caucasiana, 3, 119-126.
• Seropian, A., Bulbulashvili, N., Krammer, H. J., Thormann, J., Hein, N., Karalashvili, E., ... & Datunashvili, A. (2024). Picking pearls from the Silk Road: Insights into the spider (Arachnida, Araneae) diversity in Georgia from the Caucasus Barcode of Life (CaBOL) project. Part III. Caucasiana, 3, 89-118.
• Iskandarova, T., Japarashvili, S., Bulbulashvili, N., & Seropian, A. (2024). The first record of Ripiphoridae (Coleoptera, Cucujiformia, Tenebrionoidea) family in Georgia. Caucasiana, 3, 19-23.
• Seropian, A., & Bulbulashvili, N. (2024). The first record of Hemineura wittmeri Badonnel, 1981 (Psocodea, Psocomorpha, Elipsocidae) from the Caucasus. Caucasiana, 3, 1-4.
• Metreveli, V., Kreft, H., & Gavashelishvili, A. (2024). Environmental covariates of chestnut blight (Cryphonectria parasitica) in Georgia (Caucasus). Forest Ecology and Management, 569, 122153.
• Hwang, B. C., Giardina, C. P., Adu-Bredu, S., Barrios-Garcia, M. N., Calvo-Alvarado, J. C., Dargie, G. C., ... & Metcalfe, D. B. (2024). The impact of insect herbivory on biogeochemical cycling in broadleaved forests varies with temperature. Nature Communications, 15(1), 6011.
• Augustijnen, H., Bätscher, L., Cesanek, M., Chkhartishvili, T., Dincă, V., Iankoshvili, G., ... & Lucek, K. (2024). A macroevolutionary role for chromosomal fusion and fission in Erebia butterflies. Science Advances, 10(16), eadl0989.
• Solak, H. M., Kreisinger, J., Čížková, D., Sezgin, E., Schmiedová, L., Murtskhvaladze, M., ... & Yanchukov, A. (2024). Altitude shapes gut microbiome composition, accounting for diet, thyroid hormone levels, and host genetics in a subterranean blind mole rat. Frontiers in Microbiology, 15, 1476845.
• BUDAGASHVILI, N., & JAVAKHISHVILI, Z. The birds of Georgia: an updated checklist using citizen science platforms. Sandgrouse 46: 1-29
• Fair, J. M., Al-Hmoud, N., Alrwashdeh, M. M., Bartlow, A. W., Balkhamishvili, S., Daraselia, I., ... & Owen, J. (2024). Transboundary determinants of avian zoonotic infectious diseases: challenges for strengthening research capacity and connecting surveillance networks. Frontiers in Microbiology, 15, 1341842.
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• Iankoshvili, G., Tsikolia, N., Barateli, N., Gabelaia, M., Seropian, A., & Tarkhnishvili, D. (2023). Two-headed parthenogenetic lizard embryo from southern Georgia. Caucasiana, 2, 199-203.
• Ninua, L., Tarkhnishvili, D., & Anderson, C. L. (2023). Genetic structure of Ponto‐Caspian trout populations shows gene flow among river drainages and supports resident Salmo rizeensis as a genetically distinct taxon. Ecology and Evolution, 13(7), e10335.
• Tarkhnishvili, D., & Iankoshvili, G. (2023). The farther, the closer: geographic proximity and niche overlap versus genetic divergence in Caucasian rock lizards. Biological Journal of the Linnean Society, blad034.
• Kropachev, I., Tarkhnishvili, D., Murtskhvaladze, M., & Galoyan, E. (2023). Living apart together: Morphological, spatial, and genetic differentiation of three sympatric rock lizard species (Lacertidae: Darevskia) of the Caucasus. Zoologischer Anzeiger, 303, 71-79.
• Tarkhnishvili, D., Yanchukov, A., & Böhne, A. (2023). Advantages, limitations, and evolutionary constraints of asexual reproduction: An empirical approach. Frontiers in Ecology and Evolution, 11, 1184306.
• Beridze, T., Japoshvili, B., Edisherashvili, T., Anderson, C., & Mumladze, L. (2023). Fish diversity assessed by eDNA detection methods in the Rioni River. Metabarcoding and Metagenomics, 7, e96780.
• Gavashelishvili, A., Chukhua, M., Sakhltkhutsishvili, K., Koptekin, D., & Somel, M. (2023). The time and place of origin of South Caucasian languages: insights into past human societies, ecosystems and human population genetics. Scientific Reports, 13(1), 21133.
• Metreveli, V., Kreft, H., Akobia, I., Janiashvili, Z., Nonashvili, Z., Dzadzamia, L., ... & Gavashelishvili, A. (2023). Potential Distribution and Suitable Habitat for Chestnut (Castanea sativa). Forests, 14(10), 2076.
• Bazzicalupo, E., Ratkiewicz, M., Seryodkin, I. V., Okhlopkov, I., Galsandorj, N., Yarovenko, Y. A., ... & Godoy, J. A. (2023). Genome‐environment association analyses reveal geographically restricted adaptive divergence across the range of the widespread Eurasian carnivore Lynx lynx (Linnaeus, 1758). Evolutionary Applications.
• Koptekin, D., Yüncü, E., Rodríguez-Varela, R., Altınışık, N. E., Psonis, N., Kashuba, N., ... & Somel, M. (2023). Spatial and temporal heterogeneity in human mobility patterns in Holocene Southwest Asia and the East Mediterranean. Current Biology, 33(1), 41-57.
• Paposhvili, N., Morkunas, J., Ninua, L., Beridze, T., Kerdikoshvili, N., Dekanoidze, D., ... & Gavashelishvili, A. (2023). Genetic connectivity between Caucasian and Northern Velvet Scoter Melanitta fusca populations and its importance for the long-term survival of the species in the Caucasus. Wildfowl, 250-261.
• Japarashvili, S., Bulbulashvili, N., Seropian, A., Chkhartishvili, T., Iankoshvili, G., Chitadze, B., ... & Memishishi, A. (2023). New and well-forgotten: DNA-assisted records of two beetle (Insecta, Coleoptera) species new for the fauna of Georgia, with an update on the distribution of some other little-studied taxa. Caucasiana, 2, 63-70.
• Seropian, A., Arsenashvili, E., Bulbulashvili, N., Shubashishvili, A., Iankoshvili, G., Todua, M., ... & Rulik, B. (2023). Into the unknown: the first barcode-assisted checklist of Psocoptera (Insecta, Psocodea) of Georgia with a census on country species richness. ZooKeys, 1168, 77.
• Augustijnen, H., Baetscher, L., Cesanek, M., Chkhartishvili, T., Dincă, V., Iankoshvili, G., ... & de Vos, J. (2023). A macroevolutionary role for chromosomal fusion and fission in Erebia butterflies. bioRxiv.  doi, 10(2023.01), 16-524200.
• Seropian, A., Otto, S., & Bulbulashvili, N. (2023). Picking pearls from the Silk Road: Insights into the spider (Arthropoda, Araneae) diversity in Georgia from the CaBOL project. Part I. Caucasiana, 2, 143-159.
• Seropian, A., Bulbulashvili, N., Otto, S., Krammer, H. J., Kachlishvili, N., & Datunashvili, A. (2023). Picking pearls from the Silk Road: Insights into the spider (Arachnida, Araneae) diversity in Georgia from the Caucasus Barcode of Life project. Part II. Caucasiana, 2, 231-297.
• Arsenashvili, E., & Seropian, A. (2023). The first barcode-assisted record of Tropidopola turanica Uvarov, 1926 (Orthoptera, Acrididae) from Georgia. Caucasiana, 2, 109-111.
• Seropian, A., Arsenashvili, E., Bulbulashvili, N., Japaridze, L. G., Zukakishvili, A., Japarashvili, S., ... & Mumladze, L. (2023). The first DNA-assisted record of Hemelytroblatta livida (Blattodea: Corydiidae) from Georgia with notes on Corydiinae species composition in the Caucasus. Caucasiana, 2, 71-75.
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• Stalling, T., & Seropian, A. (2022). The Common Ant Cricket Myrmecophilus acervorum (Panzer,[1799]), new to the fauna of Georgia, southern Caucasus ecoregion (Orthoptera: Myrmecophilidae), with additional data on Myrmecophilus hirticaudus Fischer von Waldheim, 1846. Caucasiana, 1, 21-23.
• Arsenashvili, E., Iankoshvili, G., Japarashvili, S., Bulbulashvili, N., & Seropian, A. (2022). Discovery of Hemilepistus (Hemilepistus) klugii (Brandt, 1833)(Isopoda, Oniscidea) in the Republic of Georgia with an overview of the ecology and biology of the subgenus. Caucasiana, 1, 13-19.
• Japarashvili, S., Bulbulashvili, N., & Seropian, A. (2022). First record of Palorus subdepressus (Wollaston, 1864)(Coleoptera: Tenebronidae) from Georgia (Sakartvelo). Caucasiana, 1, 51-52.
• Tarkhnishvili, D., Hille, A., Waller, T., Todua, M., Murtskhvaladze, M., & Böhme, W. (2022). Morphological trends and genetic divergence in anacondas, genus Eunectes Wagler, 1830 (Serpentes: Boidae). Amphibia-Reptilia, 1(aop), 1-15.
• Barateli, N., Tarkhnishvili, D., Iankoshvili, G., & Kokiashvili, L. (2022). Reproductive effort of unisexual and bisexual rock lizards (genus Darevskia). Zoologischer Anzeiger, 301, 196-204.
• Yanchukov, A., Tarkhnishvili, D., Erdolu, M., Şahin, M. K., Candan, K., Murtskhvaladze, M., ... & Galoyan, E. (2022). Precise paternal ancestry of hybrid unisexual ZW lizards (genus Darevskia: Lacertidae: Squamata) revealed by Z-linked genomic markers. Biological Journal of the Linnean Society, 136(2), 293-305.
• Koptekin, D., Yüncü, E., Rodríguez-Varela, R., Altınışık, N. E., Psonis, N., Kashuba, N., ... & Somel, M. (2022). Spatial and temporal heterogeneity in human mobility patterns in Holocene Southwest Asia and the East Mediterranean. Current Biology.
• Darul, R., Gavashelishvili, A., Saveljev, A. P., Seryodkin, I. V., Linnell, J. D., Okarma, H., ... & Schmidt, K. (2022). Coat Polymorphism in Eurasian Lynx: Adaptation to Environment or Phylogeographic Legacy?. Journal of Mammalian Evolution, 29(1), 51-62.
• Bazzicalupo, E., Lucena‐Perez, M., Kleinman‐Ruiz, D., Pavlov, A., Trajçe, A., Hoxha, B., ... & Godoy, J. A. (2022). History, demography and genetic status of Balkan and Caucasian Lynx lynx (Linnaeus, 1758) populations revealed by genome‐wide variation. Diversity and Distributions, 28(1), 65-82.
• Beridze, T., Scheele, F., Edisherashvili, T., & Anderson, C. (2022). Rare but Not Gone: A Relict Population of the Black Sea Ship Sturgeon Acipenser nudiventris Persists in the Rioni River, Georgia. Diversity, 14(12), 1102.
• Iankoshvili, G., & Tarkhnishvili, D. (2021). Distribution of snakes (Reptilia: Serpentes) in Georgia: Social media networks help to improve scientific knowledge. Zoology in the Middle East, 67(3), 228-239.
• Barateli, N., Tarkhnishvili, D., Iankoshvili, G., Kokiashvili, L., Dvali, N., & Janiashvili, Z. (2021). Fine-scale analysis of habitat occupancy by Kura lizard (Darevskia portschinskii) and its daughter parthenogenetic form (Darevskia dahli). Herpetozoa, 34, 71.
• Gavashelishvili, A., Yanchukov, A., Tarkhnishvili, D., Murtskhvaladze, M., Akhvlediani, I., & Kazancı, C. (2021). Landscape genetics and the genetic legacy of Upper Palaeolithic and Mesolithic hunter-gatherers in the modern Caucasus. Scientific reports, 11(1), 1-12.
• Bazzicalupo, E., Lucena‐Perez, M., Kleinman‐Ruiz, D., Pavlov, A., Trajçe, A., Hoxha, B., ... & Godoy, J. A. (2021). History, demography and genetic status of Balkan and Caucasian Lynx lynx (Linnaeus, 1758) populations revealed by genome‐wide variation. Diversity and Distributions.
• Zafar-ul Islam, M., Gavashelishvili, A., Kokiashvili, L., al Boug, A., & Shehri, A. A. (2021). Modeling the distribution and movement intensity of the Arabian Leopard Panthera pardus nimr (Mammalia: Felidae). Zoology in the Middle East, 67(2), 106-118.
• Darul, R., Gavashelishvili, A., Saveljev, A. P., Seryodkin, I. V., Linnell, J. D., Okarma, H., ... & Schmidt, K. (2021). Coat Polymorphism in Eurasian Lynx: Adaptation to Environment or Phylogeographic Legacy?. Journal of Mammalian Evolution, 1-12.
• Beridze, T., Edisherashvili, T., Anderson, C., & Scheele, F. (2021). Ship sturgeon rediscovered in the Rioni River in Georgia. Oryx, 55(1), 9-9.
• Beridze, T., Boscari, E., Scheele, F., Edisherashvili, T., Anderson, C., & Congiu, L. (2021). Interspecific hybridization in natural sturgeon populations of the Eastern Black Sea: the consequence of drastic population decline?. Conservation Genetics, 1-6.

2008-2020 წწ.

1. Tarkhnishvili, D., Gabelaia, M., & Adriaens, D. (2020). Phenotypic divergence, convergence and evolution of Caucasian rock lizards (Darevskia). Biological Journal of the Linnean Society, 130(1), 142-155.
2. Tarkhnishvili, D., Barateli, N., Murtskhvaladze, M., & Iankoshvili, G. (2020). Estimating phenotypic heritability of sexual and unisexually reproducing rock lizards (genus Darevskia). Zoologischer Anzeiger, 285, 105-113.
3. Murtskhvaladze, M., Tarkhnishvili, D., Anderson, C. L., & Kotorashvili, A. (2020). Phylogeny of caucasian rock lizards (Darevskia) and other true lizards based on mitogenome analysis: Optimisation of the algorithms and gene selection. Plos one, 15(6), e0233680.
4. Tarkhnishvili, D., Yanchukov, A., Şahin, M. K., Gabelaia, M., Murtskhvaladze, M., Candan, K., ... & Ilgaz, Ç. (2020). Genotypic similarities among the parthenogenetic Darevskia rock lizards with different hybrid origins. BMC Evolutionary Biology, 20(1), 1-25.
5. Galoyan, E., Moskalenko, V., Gabelaia, M., Tarkhnishvili, D., Spangenberg, V., Chamkina, A., & Arakelyan, M. (2020). Syntopy of two species of rock lizards (Darevskia raddei and Darevskia portschinskii) may not lead to hybridization between them. Zoologischer Anzeiger, 288, 43-52.
6. Shakarashvili, M., Kopaliani, N., Gurielidze, Z., Dekanoidze, D., Ninua, L., & Tarkhnishvili, D. (2020). Population genetic structure and dispersal patterns of grey wolves (Canis lupus) and golden jackals (Canis aureus) in Georgia, the Caucasus. Journal of Zoology, 312(4), 227-238.
7. Zafar-ul Islam, M., Volmer, R., al Boug, A., Shehri, A. A., & Gavashelishvili, A. (2020). Modelling the effect of competition for prey and poaching on the population of the Arabian Leopard, Panthera pardus nimr, in Saudi Arabia (Mammalia: Felidae). Zoology in the Middle East, 66(2), 95-106.
8. Islam, M. Z., Gavashelishvili, A., Kokiashvili, L., al Boug, A., as Shehri, A., Peterson, A. T., & García, D. J. (2020). Modeling the distribution and movement intensity of Arabian Leopard Panthera pardus nimr in the Arabian Peninsula. Authorea Preprints.
9. Kuemmerle, T., Bluhm, H., Ghoddousi, A., Arakelyan, M., Askerov, E., Bleyhl, B., Gavashelishvili, A., ... & Manvelyan, K. (2020). Identifying priority areas for restoring mountain ungulates in the Caucasus ecoregion. Conservation Science and Practice, 2(11), e276.
10. Mikeladze, G., Gavashelishvili, A., Akobia, I., & Metreveli, V. (2020). Estimation of forest cover change using Sentinel-2 multi-spectral imagery in Georgia (the Caucasus). iForest-Biogeosciences and Forestry, 13(4), 329.
11. Chaladze, G. (2020). Estimation of Ground Dwelling Arthropod Densities Using Pitfall Traps: Individual Based Simulations. Evolutionary Biology, 5(1), 1-5.
12. Zerbe, S., Pieretti, L., Elsen, S., Asanidze, Z., Asanidze, I., & Mumladze, L. (2020). Forest Restoration Potential in a Deforested Mountain Area: An Ecosociological Approach towards Sustainability. Forest Science, 66(3), 326-336.
13. Mumladze, L., Japoshvili, B., & Anderson, E. P. (2020). Faunal biodiversity research in the Republic of Georgia: a short review of trends, gaps, and needs in the Caucasus biodiversity hotspot. Biologia, 75(9), 1385-1397.
14. Thormann J, Ahrens D, Anderson C, Astrin JJ, Mumladze L , Rulik B, Tarkhnishvili D, Espeland M, Geiger M, Hein N, Iankoshvili G, Karalashvili E, Mengual X, Morkel C, Neiber MT, Peters RS, Reimann A, Ssymank A , Wesener T, Ziegler J, Misof B. 2019. A prelude to the Caucasus Barcode of Life Platform (CaBOL): Biodiversity Days in Georgia in 2018 and 2019. Bonn Zoological Bulletin 68 (2): 275–296.
15. Barjadze S., Asanidze Z., Gavashelishvili A., Soto-Adams FN. 2019. The hypogean invertebrate fauna of Georgia (Caucasus). Zoology in the Middle East 65, 1-10.
16. Bleyhl B, Arakelyan M, Askerov E, Bluhm H, Gavashelishvili A, Ghasabian M, Ghoddousi A, Heidelberg A, Khorozyan I, Malkhasyan A, Manvelyan K, Masoud M, Moqanaki EM, Radeloff VC, Soofi M, Weinberg P, Zazanashvili N, Kuemmerle T. 2019. Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Diversity and Distributions 25: 129-141.
17. H Pretzsch, M Steckel, M Heym, P Biber, C Ammer, M Ehbrecht, K Bielak, F Bravo, C Ordóñez, C Collet, F Vast, L Drössler, G Brazaitis, K Godvod, A Jansons, J de-Dios-García, M Löf, J Aldea, N Korboulewsky, DOJ Reventlow, A Nothdurft, M Engel, M Pach, J Skrzyszewski, M Pardos, Quentin Ponette, R Sitko, M Fabrika, M Svoboda, J Černý, B Wolff, R Ruíz-Peinado, M del Río. 2019. Stand growth and structure of mixed-species and monospecific stands of Scots pine (Pinus sylvestris L.) and oak (Q. robur L., Quercus petraea (MATT.) LIEBL.) analysed along a productivity gradient through Europe. European Journal of Forest Research 
18. Géraud de Streel, C Collet, I Barbeito, K Bielak, Andres Bravo-Oviedo, G Brazaitis, L Coll, L Drössler, D Forrester, M Heym, M Löf, M Pach, H Pretzsch, R Ruiz-Peinado, J Skrzyszewski, J Stankevičiūtė, M Svoboda, K Verheyen, T Zlatanov, D Bonal, Quentin Ponette. 2019. Contrasting patterns of tree species mixture effects on wood δ13C along an environmental gradient—European Journal of Forest Research. 
19. Peter Freer-Smith, Bart Muys, Michele Bozzano, Lars Drössler, Niall Farrelly, Hervé Jactel, Jaana Korhonen, Gianfranco Minotta, Maria Nijnik, Christophe Orazio. 2019. Plantation forests in Europe: challenges and opportunities. European Forest Institute 9, 1-52.
20. Gabelaia M, D Tarkhnishvili, D Adriaens. 2018. Use of three-dimensional geometric morphometrics for the identification of closely related species of Caucasian rock lizards (Lacertidae: Darevskia). Biological Journal of the Linnean Society, Volume 125, Issue 4, 26 November 2018, Pages 709–717,
21. Gavashelishvili A., YA Yarovenko, EA Babayev, GMikeladze, Z Gurielidze, D Dekanoidze, N Kerdikoshvili, L Ninua, N Paposhvili. 2018. Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus. Journal of Mammalogy, xx(x):1–13,
22. Axel Barlow, James A Cahill, Stefanie Hartmann, Christoph Theunert, Georgios Xenikoudakis, Gloria G Fortes, Johanna LA Paijmans, Gernot Rabeder, Christine Frischauf, Aurora Grandal-d’Anglade, Ana García-Vázquez, M Murtskhvaladze, Urmas Saarma, Peeter Anijalg, Tomaž Skrbinšek, Giorgio Bertorelle, Boris Gasparian, Guy Bar-Oz, Ron Pinhasi, Montgomery Slatkin, Love Dalén, Beth Shapiro, Michael Hofreiter, 2018. Partial genomic survival of cave bears in living brown bears. Nature ecology & evolution 2: 1563
23. M Nassirkhani, L Mumladze 2018. A new cave pseudoscorpion (Pseudoscorpiones: Neobisiidae) from western Georgia. Arachnology 17 (9), 496-500
24. M Murvanidze, N Todria, L Mumladze, L Kalatozishvili. 2018. Diversity of soil mite communities in different habitats of the Saskhori quarries, Georgia. Persian Journal of Acarology 7
25. Sophio Gabelashvili, Levan Mumladze, Ani Bikashvili, Pavel Sroka, Roman J Godunko, Bella Japoshvili. 2018. The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country. Turkish Journal of Zoology 42: 252-262.
26. T Kuljanishvili, L Mumladze, L Kalous, B Japoshvili 2018. Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia). Biologia 73 (1), 93-100
27. Ninua L, Tarkhnishvili D, Gvazava E. 2018. Phylogeography and taxonomic status of trout and salmon from the Ponto ‐Caspian drainages, with inferences on European Brown Trout evolution and taxonomy. – Ecology and Evolution 8: 2645-2658
28. Gabelashvili A, Mumladze M, Bikashvili A, Sroka P, Godunko RJ, Japoshvili B. 2018. The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country. Turkish J. Zool. 42: 252-262.
29. Kuljanishvili T, Mumladze L, Kalous L, Japoshvili B. 2018. Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia). Biologia, 73: 93-100.
30. Tarkhnishvili D., M Gabelaia, A Kandaurov, A Bukhnikashvili, G Iankoshvili. 2017. Isolated population of the Middle Eastern Phoenicolacerta laevis from the Georgian Black Sea Coast, and its genetic closeness to populations from southern Turkey. Zoology in the Middle East. 63, 311-315
31. Gabelaia M, D Adriaens, D Tarkhnishvili. 2017. Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger-A Journal of Comparative Zoology 268, 32-40
32. Tarkhnishvili D, M Murtskhvaladze, Cl Anderson. 2017. Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linnean Society, 121, 365–378.
33. Tarkhnishvili D, A Gavashelishvili, M Murtskhvaladze, A Latsuzbaia. 2017. Landscape Complexity in the Caucasus impedes Genetic Assimilation of Human Populations More Effectively than Language or Ethnicity. Human Biology 88, 287-300
34. Mumladze L, W Ulrich, Z Asanidze, G Japoshvili. 2017. An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Écoscience, 1-5
35. Mumladze L, M Murvanidze, M Maraun. 2017. Patterns of oribatid mite species diversity: testing the effects of elevation, area and sampling effort. Experimental and Applied Acarology, 1-18.
36. Aslan E, L Mumladze, G Japoshvili. 2017. List of leaf beetles (Coleoptera: Chrysomelidae) from Lagodekhi reserve with new records for Transcaucasia and Georgia, Zootaxa 42277 (1), 086-098
37. Japoshvili B, L Mumladze, L Murvanidze. 2017. The population of Carassius gibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iranian Journal of Fisheries Sciences 16 (2), 793-799
38. Mumladze L, Z Asanidze, F Walther, B Hausdorf. 2017. Beyond elevation: testing the climatic variability hypothesis vs Rapoport’s rule in vascular plant and snail species in the Caucasus. Biological Journal of the Linnean Society, 121, 753–763
39. Chaladze G. 2017. Taxonomy should be more photography-based, eliminating the need for a physical specimen to study morphology. Zootaxa 4247 (3), 331-331
40. Chaladze G, L Kalatozishvili, Z Janiashvili, G Bakuradze. 2017. Pterostichus neilgaimani sp. nov., a new species of ground beetles (Coleoptera: Carabidae) from a relict sacred grove in Eastern Georgia. Zootaxa 4329 (1), 91-95.
41. Gavashelishvili A, D Tarkhnishvili. 2016. Biomes and human distribution during the last ice age. Global Ecology and Biogeography, DOI: 10.1111/geb.12437 ISI IF 5.84
42. Mumladze, L., Asanidze, Z., Walther, F., Hausdorf, B. (2017). Beyond elevation: Testing the climatic variability hypothesis versus Rapoport’s rule in vascular plant and snail species in the Caucasus. Oecologia (Accepted). ISI IF 2.902
43. Chaladze G. 2016. Heterosexual Male Carriers Could Explain Persistence of Homosexuality in Men: Individual-Based Simulations of an X-Linked Inheritance Model. Archives of Sexual Behavior, 1-7. ISI IF 2.704
44. Tarkhnishvili D, M. Murtskhvaladze, CL Anderson. 2017. Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linnean Society (Accepted) ISI IF 1.984
45. Murvanidze M, L Mumladze. 2016. Annotated checklist of Georgian oribatid mites. Zootaxa 4089 (1), 1-81. ISI IF 0.994
46. Tarkhnishvili D, M Gabelaia, L Mumladze, M Murtskhvaladze. 2016. Mitochondrial phylogeny of the Darevskia saxicola complex: two highly deviant evolutionary lineages from the easternmost part of the range. The Herpetological Journal 26 (2), 175-182. ISI IF 0.808
47. Tarkhnishvili D, A. Gavashelishvili, M Murtskhvaladze (2017) Landscape Complexity in the Caucasus Impedes Genetic Assimilation of Human Populations More Effectively Than Language or Ethnicity. Human Biol. (Accepted). ISI IF 0.857
48. Japoshvili, B., Mumladze, L., & Murvanidze, L. (2017). The population of Carssiusgibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iran J Fish Sci (Accepted). ISI IF 0.393
49. Murvanidze M, L Mumladze, T Arabuli, S Barjadze, M Salakaia. 2016. Oribatida diversity in different microhabitats of Mtirala National Park. J. Acarol. Soc. Jpn 25 (Supplement 1), S35-S49. ISI IF N/A
50. Bakhtadze, N.G., Chakvetadze, N.L., Mumladze, L.J., Bakhtadze, G.I., & Tskhadaia, E.A. (2016). Karyological Data of Terrestrial Molluscs (Mollusca: Gastropoda: Pulmonata) of Georgia. Proceedings of the Institute of Zoology 12/2016; XXV: 23-27 ISI IF N/A
51. Gabelashvili, S., Bikashvili, A., Shubitidze, Z., Gioshvili, M., Pankvelashvili, E., Mumladze, L., & Japoshvili, B. (2016). Family-level diversity and distribution of macroinvertebrates in Madatapa, Khanchali, and Bughdasheni lakes in the Javakheti Plateau (South Georgia). Proceedings of the Institute of Zoology 12/2016; XXV: 117-127 ISI IF N/A
52. Mumladze, L., & Paposhvili, N. (2016). A New Addition to the Malacofauna of Georgia – Eobania Vermiculata is Replenishing its Range. Proceedings of the Institute of Zoology 12/2016; XXV:153-155 ISI IF N/A
53. Seehausen, M., Schröter, A., Mumladze, L., Grebe, B. (2016). Additional Odonata records from Georgia, southern Caucasus ecoregion, with the first record of Ischnura fountaineae (Odonata: Coenagrionidae). Notulae Odonatologicae. 8 (8), 266-283 ISI IF N/A
54. Gabelaia, M., Tarkhnishvili, D. and Murtskhvaladze, M. 2015. Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta. Amphibia-Reptilia 36: 45-54.
55. Mumladze, L., Murvanidze, M., Maraun, M. & Salakaia, M. (2015). Oribatid mite communities along an elevational gradient in Sairme gorge (Caucasus). Experimental and Applied Acarology. 66(1), 41-51.
56. Farhadinia, M. S., Farahmand, H., Gavashelishvili, A., Kaboli, M., Karami, M., Khalili, B., Montazamy, S. 2015. Molecular and craniological analysis of leopard, Panthera pardus (Carnivora: Felidae) in Iran: support for a monophyletic clade in Western Asia. Biological Journal of the Linnean Society 114: 721-736.
57. Lewis NS, Verhagen JH, Javakhishvili Z, Russell CA, Lexmond, P. et al (20 authors). 2015. Influenza A virus evolution and spatio-temporal dynamics in Eurasian Wild Birds: A phylogenetic and phylogeographic study of whole-genome sequence data. Journal of General Virology
58. Tarkhnishvili D. 2014. Rana macrocnemis. In: Grossenbacher, K. (ed.), Handbuch Amphibien & Reptilien Europas. Aula-Verlag, Germany.
59. Kopaliani, N., Shakarashvili, M., Gurielidze, Z., Qurkhuli, T. & Tarkhnishvili, D. 2014. Gene flow between wolves and shepherd dogs in the Caucasus. J. Hered. (in press).
60. Mumladze L, Cameron RAD, Pokryszko BM. 2014. Endemic land molluscs in Georgia (Caucasus): how well are they protected by existing reserves and national parks? Journal of Molluscan Studies 80: 67-73.
61. Tarkhnishvili D, Murtskhvaladze M, Gavashelishvili A. 2013. Speciation in Caucasian lizards: climatic dissimilarity of the habitats is more important than isolation time. Biological Journal of the Linnean Society, 109, 876-892.
62. Mumladze L., Tarkhnishvili D., Murtskhvaladze M. 2013. Systematics and evolutionary history of large endemic snails from the Caucasus (Helix buchii and H. goderdziana) (Helicidae) American Malacological Bulletin 31 (2), 225-234. 
63. Mumladze L, Murvanidze M, Behan-Pelletier V. 2013. Compositional patterns in Holarctic peat bog-inhabiting oribatid mite (Acari: Oribatida) communities Pedobiologia 55 (1), 41-48
64. Murvanidze M, Mumladze L, Arabuli T, Kvavadze E. 2013. Oribatid mite colonization of sand and manganeze tailing sites. Acarologia 53 (2), 203-215
65. Japoshvili B, Mumladze L, Kucuk F. 2013. Invasive Carassius Carp in Georgia: Current state of knowledge and future perspectives. Current Zoology, in press
66. Mumladze L. 2013. Shell size differences in Helix lucorum Linnaeus, 1758 (Mollusca: Gastropoda) between natural and urban environments. Turk J. Zool. 36: 1-7
67. Lewis NS, Z Javakhishvili, CA Russell, A Machablishvili, P Lexmond, JH. 2013.Avian Influenza Virus Surveillance in Wild Birds in Georgia: 2009–2011. PloS one 8 (3), e58534 2008-2012: 20
68. Tarkhnishvili, D. 2012: Evolutionary History, Habitats, Diversification, and Speciation in Caucasian Rock Lizards. Advances in Zoology Research. Owen P. Jenkins, Ed. 2, 79-120. Nova Scientific Publishers, NY.
69. Tarkhnishvili D., Gavashelishvili A., Mumladze L. 2012: Palaeoclimatic models help to understand current distribution of Caucasian forest species. Biological Journal of the Linnean Society. 105: 231-248.
70. Khorozyan I, Malkhasyan A, Murtskhvaladze M. 2012. The striped hyaena Hyaena hyaena (Hyaenidae, Carnivora) rediscovered in Armenia. Folia Zoologica 60: 253-261.
71. Murtskhvaladze M., Gurielidze Z., Kopaliani N., Tarkhnishvili D. 2012: Gene introgression between Gazella subguturrosa and G. marica: limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica 
72. Gavashelishvili A, McGrady M, Ghasabian M, Bildstein KL. 2012. Movements and habitat use by immature Cinereous Vultures (Aegypius monachus) from the Caucasus. Bird Study 59 (4): 449-462.
73. Chaladze G. 2012: Climate-based model of spatial pattern of the species richness of ants in Georgia. Journal of Insect Conservation. 
74. Murvanidze M, Mumladze L, Arabuli T, Kvavadze E. 2011. Landscape distribution of oribatid mites (Acari, Oribatida) in Kolkheti National Park (Georgia, Caucasus). Zoosymposia 6: 221-233.
75. Murvanidze M, Kvavadze E, Mumladze L, Arabuli T. 2011. Comparison of Earthworms (Lumbricidae) and Oribatid Mite (Acari, Oribatida) Communities in Natural and Urban Ecosystems. Vestnik Zoologii, 45: 16-24.
76. Pokryszko B, Cameron R, Mumladze L, Tarkhnishvili D. 2011: Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society. 102: 239-250.
77. Tarkhnishvili, D. Chaladze, G. et al., 2011. Georgian biodiversity database. www.biodiversity-georgia.net  
78. Tarkhnishvili D., Gavashelishvili A., Avaliani A., Murtskhvaladze M., Mumladze L. 2010: Unisexual rock lizard might be outcompeting its bisexual progenitors in the Caucasus. Biological Journal of the Linnean Society. 101: 447-460.
79. Murtskhvaladze M., Gavashelishvili A., Tarkhnishvili D. 2010: Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus. Molecular Ecology. 19: 1829-1841.
80. Gavashelishvili A, Javakhishvili Z. 2009: Combining Radio-Telemetry and Random Observations to Model the Habitat of Caucasian Grouse (Tetrao Mlokosiewiczi). Oryx. 44: 491-500. 
81. Tarkhnishvili D., Serbinova I., Gavashelishvili A. 2009: Modelling the range of Syrian spadefoot toad (Pelobates syriacus) with a combination of GIS-based approaches. Amphibia-Reptilia. 30: 401-412
82. Sayim, F., Baskale, E., Tarkhnishvili, D., Kaya, U. 2009. Some water chemistry parameters of the breeding habitats of the Caucasian salamander, Mertensiella caucasica, in the Western Lesser Caucasus. Comptes Rendus Biologies, Vol. 332, No. 5. (May 2009), pp. 464-469.
83. Tarkhnishvili D., Kaya U. 2009: Status and Conservation of the Caucasian Salamander (Mertensiella caucasica). In: Zazanashvili, N., Mallon, D. (eds), Status And Protection of Globally Threatened Species in the Caucasus, pp. 157-165.
84. Gavashelishvili A. 2009: GIS-based Habitat Modeling of Mountain Ungulate species in the Caucasus Hotspot. In: Zazanashvili, N., Mallon, D. (eds), Status And Protection of Globally Threatened Species in the Caucasus, pp. 74-82
85. Sinsch U., Schneider H. & Tarkhnishvili D. 2008 Bufo bufo Superspecies – Erdkröten-Artenkreis. PP. 191-335 In: Grossenbacher, K. (ed.), Handbuch Amphibien & Reptilien Europas. Aula-Verlag, Germany
86. Mumladze L., Tarkhnishvili D., Pokryszko B. 2008: A new species of the genus Helix from south-western Georgia. Journal of Conchology 39: 483-485. 
87. Kvavadze E, Rukhadze L, Nikolaishvili V, Mumladze L. 2008. Botanical and zoological remains from an early medieval grave at Tsitsamuri, Georgia. Vegetation History and Archaeobotany 17 (1), 217-224
88. Angus RB, Galián J, Wrase DW, Chaladze G. 2008. The western Palaearctic species of the Pterostichus nigrita (Paykull) complex, with the description of a new species from Spain and a new subspecies of P. nigrita from Anatolia (Coleoptera, Carabidae). Nouv Rev Entomol 25, 297-316
89. Gavashelishvili A., Lukarevsky V. 2008: Modelling the habitat requirements of the leopard Panthera pardus in West and Central Asia. Journal of Applied Ecology 45: 579-588.
90. Poulakakis N, Antoniou A, Mantziou G, Parmakelis A, Skartsi T, Vasilakis D, Elorriaga J, De La Puente J, Gavashelishvili A, Ghasabyan M, Katzner T, Mcgrady M, Batbayar N, Fuller M, Natsagdorj T, 2008. Population structure, diversity, and phylogeography in the near‐threatened Eurasian black vultures Aegypius monachus (Falconiformes; Accipitridae) in Europe: insights from microsatellite and mitochondrial DNA variation. Biological Journal of the Linnean Society, 95: 859-872.
91. Tarkhnishvili D., Kaya U., Gavashelishvili A., Serbinova I. 2008: Ecological Divergence between two Evolutionary Lineages of the Caucasian Salamander: Evidence from the GIS analysis. Herpetological Journal. 18: 155-163

მსხვილი ძუძუმწოვრების ეკოლოგიის პროგრამა 

2021-2025წწ.

• Stefanović M , Bogdanowicz W,  Adavoudi R ,  Martínez-Sosa F , Doan K, Alejandro Flores-Manzanero A,  Srinivas Y, Ovidiu C.. , Ćirović D ,  D’Amico G. ,  Djan M,  Giannatos G, Jennifer Hatlauf J, Hayrapetyan V., M,  Homel K.,  Hulva P., Ionică A., Vikramsinh Jhala Y. ,  Juránková J. ,  Kaboli M.,  Khosravi R., Kopaliani N., Kowalczyk R.,  Krofel M., Lanszki J , Lapini L., Lymberakis P., Männil P, Markov G.,  Mihalca A., Miliou A., Modrý D.,  Molchan V.,  Ostrowski S. ,  Pakeltytė G.,  Ruņģis D,  Šnjegota D, Szabó L. ,  Tryfonopoulos G.A., Tsingarska E., Volokh A., Wójcik J., Pilot M. 2025. Range-wide phylogeography of the golden jackals (Canis aureus) reveals multiple sources of recent spatial expansion and admixture with dogs at the expansion front. Accepted by Biological Conservation.
• Ivanchikova, Julia & Tregenza, Nick & Popov, Dimitar & Meshkova, Galina & Marian, Paiu & Timofte, Costin & Öztürk, Ayaka & Tonay, M. & Dede, Ayhan & Özsandıkçı, Uğur & Kopaliani, Natia & Dekanoidze, Davit & Gurielidze, Zurab & Vishnyakova, Karina & Hammond, Philip & Gol'din, Pavel. (2024). Seasonal and diel patterns in Black Sea harbour porpoise acoustic activity in 2020–2022. Ecology and Evolution. 14. 10.1002/ece3.70182.
• Ninua L., Tarkhnishvili D., Anderson C.L. (2023) Genetic structure of Ponto-Caspian trout populations shows gene flow among river drainages and supports resident Salmo rizeensis as a genetically distinct taxon. Ecology & Evolution. 2023 Jul 24;13(7):e10335. doi: 10.1002/ece3.10335. eCollection 2023 Jul.
  Klink AC, Rula O, Sushko M, Bezymennyi M, Mezinov O, Gaidash O, Bai X, Stegniy A, Sapachova M, Datsenko R, Skorokhod S, Nedosekov V, Hill NJ, Ninua L, Kovalenko G, Ducluzeau AL, Mezhenskyi A, Buttler J, Drown DM, Causey D, Stegniy B, Gerilovych A, Bortz E, Muzyka D. Discovery of Avian Paramyxoviruses APMV-1 and APMV-6 in Shorebirds and Waterfowl in Southern Ukraine. Viruses. 2023 Mar 8;15(3):699. doi: 10.3390/v15030699. PMID: 36992408; PMCID: PMC10058161.

• Šnjegota D. ,  Niedziałkowska M. , Stronen A.V. , Borowik T.,  Plis K.,  Arakelyan M.,  Ćirović  D , Gabriel Danila G. ,  Djan M, Ghazaryan A., Gurielidze Z. , Hayrapetyan T.,  Hegyeli Z., Karamanlidis A.A. , Kopaliani N. ,  Kusak ., Politov D. , Talala M. , Tsingarska E ,  Jędrzejewska B. The role of the Caucasus, Carpathian, and Dinaric-Balkan regions in preserving wolf genetic diversity. Mammalian Biology.

• Klink, A.C.; Rula, O.; Sushko, M.; Bezymennyi, M.; Mezinov, O.; Gaidash, O.; Bai, X.; Stegniy, A.; Sushko, M.; Datsenko, R.; Skorohod, S.; Nedosekov, V.; Hill, N.; Ninua, L.; Kovalenko, G.; Ducluzeau, A.L.; Mezhenskiy, A.; Drown, D.M.; Causey, D.; Stegniy, B.; Gerilovych, A.; Bortz, E.; Muzyka, D. Discovery of Avian Paramyxoviruses APMV-1 and APMV-6 in Shorebirds and Waterfowl in Southern Ukraine . Preprints 2022, 2022060322 (doi: 10.20944/preprints202206.0322.v1).

• Pilot, M., Moura, A. E., Okhlopkov, I. M., Mamaev, N. V., Manaseryan, N. H., Hayrapetyan, V., Kopaliani, N., Tsingarska, E., Alagaili, A. N., Mohammed, O. B., Ostrander, E. A., & Bogdanowicz, W. (2021). Human-modified canids in human-modified landscapes: The evolutionary consequences of hybridization for grey wolves and free-ranging domestic dogs. Evolutionary Applications, 14, 2433–2456. https://doi.org/10.1111/eva.13257
• Bazzicalupo E., Lucena-Perez M., Kleinman-Ruiz D.,  Pavlov A., Trajçe A.,  Hoxha B.,   Sanaja B.,  Zurab Gurielidze Z., Kerdikoshvili N., Mamuchadze J., Yarovenko Y.,  Akkiev M., Ratkiewicz M., Alexander P., Saveljev A.P.,  Melovski D., Gavashelishvili A., Schmidt K.,  Godoy J.A. 2021. History, demography and genetic status of Balkan and Caucasian Lynx lynx (Linnaeus, 1758) populations revealed by genome-wide variation. Diversity and Distributions. 2021;00:1–18.

2006-2020 წწ.

1. Shakarashvili N. Kopaliani Z. Gurielidze  D. Dekanoidze  L. Ninua  D. Tarkhnishvil. 2020.  Population genetic structure and dispersal patterns of grey wolf (Canis lupus) and golden jackal (Canis aureus) in Georgia, the Caucasus—Journal of Zoology.
2. ნათია კოპალიანი, ზურაბ გურიელიძე, დავით დეკანოიძე, ივანე დარასელია, ნანა დევიძე. 2020. მართვის გეგმა ზურმუხტის ქსელის ტერიტორიისთვის სვანეთი 1 (Svaneti 1 – GE0000012). ილიას სახელმწიფო უნივერსიტეტი. მომზადდა გარემოს დაცვის და სოფლის მეურნეობის სამინისტროს ხელშეწყობით. თბილისი,2020
3. ნათია კოპალიანი, ზურაბ გურიელიძე, დავით დეკანოიძე, ივანე დარასელია, ნანა დევიძე. 2020. მართვის გეგმა ზურმუხტის ქსელის ტერიტორიისთვის სვანეთი 2 (Svaneti 2 – GE0000045). ილიას სახელმწიფო უნივერსიტეტი. მომზადდა გარემოს დაცვის და სოფლის მეურნეობის სამინისტროს ხელშეწყობით. თბილისი,2020
4. Pilot M, Moura A., Okhlopkov I, Mamaev N, Alagaili A., Mohammed O., Eduard Yavruyan E., Hayrapetyan V., Kopaliani N, Tsingarska E., Krofel M, Skoglund P. & Bogdanowicz W. 2019. Global Phylogeographic and Admixture Patterns in Grey Wolves and Genetic Legacy of An Ancient Siberian Lineage. Scientific Reports. (2019) 9: 1732Ninua L, Tarkhnishvili D, Gvazava E. Phylogeography and taxonomic status of trout and salmon from the Ponto-Caspian drainage, with the inference on the European Brown Trout evolution and taxonomy. Ecology and Evolution 2018 Mar; 8(5): 2645–2658
5. Divya Venkatesh , Marjolein J. Poen, Theo M. Bestebroer , Rachel D. Scheuer, Oanh Vuong , Mzia Chkhaidze , Anna Machablishvili , Jimsher Mamuchadze , Levan Ninua, Nadia B. Fedorova , Rebecca A. Halpin , Xudong Lin , Amy Ransier , Timothy B Stockwell , David E. Wentworth, Divya Kriti, Jayeeta Dutta , Harm van Bakel , Anita Puranik , Marek J Slomka , Steve Essen , Ian H. Brown, Ron A.M. Fouchier , Nicola S. Lewis. Avian influenza viruses in wild birds: virus evolution in a multi-host ecosystem. Journal of Virology, 16 May 2018
6. Alexander Gavashelishvili, Yuriy A Yarovenko, Elmar A. Babayev, Giorgi Mikeladze, Zurab Gurielidze, Davit Dekanoidze, Niko Kerdikoshvili, Levan Ninua, Nika Paposhvili. Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus. Journal of Mammalogy, xx(x):1–13, 2018 
7. Kopaliani N, Gurielidze Z, Ninua L. 2017.Records of anomalously white harbour porpoises and atypical pigmented short-beaked common dolphins in the Georgian Black Sea waters. Journal of Black Sea/ Mediterranean Environment http://www.blackmeditjournal.org/index.php/component/k2/item/575
8. Shvelidze T (2016). First Record of Oithona davisae Ferrari and Orsi, 1984 (Copepoda, Cyclopoida) and Seasonal Variations in the Georgian Black Sea Coast. Acad. J. Sci. Res. 4(11): 424-428.
9. Rutkowski R, Krofel M, Giannatos G, Ćirović D, Männil P, Volokh A., Lanszki J, Heltai M, Ovidiu C. Banea, Hayrapetyan V, Kopaliani N, Miliou A. Tryfonopoulos G., Lymberakis P, Penezić A., Pakeltytė G., Suchecka E., Bogdanowicz W. 2015. A European Concern? Genetic Structure and Expansion of Golden Jackals (Canis aureus) in Europe and the Caucasus. PLOS ONE. November 5, 2015
10. Mallon D., Weinberg P. and N. Kopaliani. Status of the Prey Species of the Leopard in the Caucasus. Cat News Special Issue. No 2. IUCN. Species Survival Commission.Cat Specialis Group. Stampfli Publikationen AG, Bern,Switzerland. 2007.
11. Lukarevsky V., Akkiev M., Askerov E., AgiliA., Can E., Gurielidze Z, Kudaktin A., Malkhasyan A and Yarovenko Y. Status of the Leopard in the Caucasus. Cat News Special Issue. No 2. IUCN. Species Survival Commission.Cat Specialis Group. Stampfli Publikationen AG, Bern, Switzerland. 2007.
12. Kopaliani N., Gurielidze Z. Status of Turs in Georgia and Conservation Action Plan. Status and Protection of Globally Threatened Species in the Caucasus. CEPF /WWF. 2009.
13. Murtskhvaladze M., Gurielidze Z., Kopaliani N., Tarkhnishvili D. Gene introgression between Gazella subguturrosa and G. marica: limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica. 4, 383-386. 2012.
14. Bogdanowicz W, Dabrowski M, Hayrapetyan V, Kopaliani N, Pilot M, Tsingarska E, Yavruyan E. Genetic variability of the grey wolf Canis lupus in the Caucasus in comparison with Europe and the Middle East: Distinct or intermediary population? PLOS ONE. 2014
15. Kopaliani N, Shakarashvili M, Gurielidze Z, Qurkhuli T and Tarkhnishvili D. Hybridization of wolf and dog in Georgia (Caucasus): introgression of wolf alleles into shepherd dog population. The Journal of Heredity. 2014.
16. ზ. გურიელიძე, ნ. კოპალიანი, ნ. დევიძე, მ. შაქარაშვილი, შავი ზღვის ბიომრავალფეროვნება. სიტუაციის ანალიზი (ქართულად და ინგლისურად). GIZ. 96 გვ. 2012.

მაღალმთის ეკოსისტემების კვლევის პროგრამა 

2021-2025წწ.

• Jolokhava, T., Abdaladze, O., Bakhia, A., Asanidze, Z., Ekhvaia, J., & Kikvidze, Z. (2025): Soil pH as a strong driver of plant species distribution in alpine-nival ecotone of the Central Caucasus. Hacquetia. https://doi.org/10.3986/hacq-2025-0022 ISI IF = 0.7
• Bussmann, RW; Paniagua Zambrana, NY; Kikvidze, Z; Sikharulidze, S; Kikodze, D; Tchelidze, D; Khutsishvili, M; Batsatsashvili, K. (2025): Dioscorides in the Caucasus: Medicinal Plants and Fungi of Sakartvelo (Republic of Georgia), Caucasus. In: Oberhelman, SM (ed): Manuscripts, Plants, Remedies in the Mediterranean Traditions: Studies across Disciplines for Alain Touwaide. vol. 2.  Plants. Botany, Gardens, “Materia medica”, Ethnopharmacology. DeGruyter/Brill, Berlin, pp. 683 - 707. ISI IF = 5.4
• Kikvidze, Mosulishvili M., Ckhutiashvili G., Toriashvili N., Maisaia I., McAlvy A., Power A. G., Paniagua Zambrana NY., Bussmann R. (2025): From field to the edge of memory: the ethnobotany and conservation of indigenous wheats in Georgia, the Caucasus. In: Ethnobotany of the Caucasus, Springer. ISI IF = 1.5
• Tedoradze G., Bakhia A., Gugutishvili N., Khuskivadze D., Mikeladze G., Kikvidze Z. (2025): Discovery of a new non-native aquatic plant in the Kolkheti Lowland, Georgia: implications for biodiversity. BioInvasions Records, Vol. 14 (4), pp. 871–882. ISI IF = 1.4
• Rusishvili N., Jalabadze M., Mosulishvili M., Batiuk S., Boaretto E., Martkophlishvili I., Meladze N., Lordkipanidze D. (2025): Kvemo Kartli (Southeast Georgia): An Independent Center of Bread Wheat Origin. In PNAS of USA. ISI IF = 9.1
• Dullinger, Chytrý K., Gattringer A., Hausharter J., Helm N., Hülber K., Moser D., Abdaladze O., Andrews Ch., Barni H., Barolin D., Benito-Alonso J-L^., von Büren R. S., Carbognani M., Carnicero P., Di Cecco V., Choler P., Dick J.,  Fernández-Calzado R., De Frenne P., García Lino M. C., Ghosn D., Gigauri Kh., Graae B. J., Hilpold A., Jiménez  J. J., Kagerl K., Kanka R., Kollár J., Lamprecht A., Lembrechts J., Lenoir J., Lorite J., Nyktas P., Pușcaș M., Randin C. F., Rixen Ch., Rossi G., Saccone P., Ursu T. M., Vanneste T., Varricchione M., Vittoz P., Winkler M., Wipf  S., Pauli H., Wessely J. (2026): High-elevation specialists increasingly disappear from European mountaintop floras. Science. Status of the article: In press, Under review. ISI IF = 49.7
• Vanneste T., Pauli H., Dullinger S., Graae B., Abdaladze O., Benito Alonso J. L. et al. (2026): Widespread shrubification on European mountain summits. Global Change Biology. Submission ID a4e834fe-3d77-4121-90d3-82b86c2570f5. ISI IF = 12.1
• Tigishvili, K., Kupradze, I. ., Jorjadze, A., Batsatsashvili, K., Kvelashvili, A. (2025): Cryptogam diversity associated with xerophilous shrubs in semiarid habitats of Georgia, South Caucasus. Bulletin of the Georgian National Academy of Sciences, 19(193), 3, 98–105. Retrieved from http://science.org.ge/bnas/vol-19-3.html ISI IF = 0.12
• Knollová I., Chytrý M., Bruelheide H., Dullinger S., Jandt U., Bernhardt-Römermann M., Biurrun I., de Bello F., Glaser M., Hennekens S., Jansen F., Jiménez-Alfaro B., Kadaš D., Kaplan E., Klinkovská K., Lenzner B., Pauli H., Gaia Sperandii M., Verheyen K., Winkler M., Abdaladze O. … (2024): ReSurveyEurope: A database of resurveyed vegetation plots in Europe. J Veg Sci. 35: e13235; DOI: 10.1111/jvs.13235 https://doi.org/10.1111/jvs.13235  ISI IF = 2.2
• Ekhvaia Zh., Abdaladze O., Jolokhava T., Bakhia A. (2024): Leaf Shape and size variability of Dryas octopetala  in the Central Great Cauasus (the Kazbegi region, Georgia). Botanica Serbica, 48 (2): 151–162, DOI: https://doi.org/10.2298/BOTSERB2402151E. ISI IF = 0.9
• Silakadze N., Mosulishvili M., Borsch T., Kilian N. (2024): The polyphyletic Caucasus-centered Campanula Scapiflorae (Campanulaceae) revisited with a newly circumscribed C. sect. Tridentatae for its core clade. PhytoKeys, 243, pp. 149-184. https://phytokeys.pensoft.net/article/120908. ISI IF = 1.4
• Sadunishvili T., Maisaia I., Sikharulidze Sh., Bussmann R.W., Paniagua Zambrana N.Y., Batsatsashvili K., Darchidze T.Foxtail millet (Setaria italica (L.) P. Beauv.) culture in Georgia, South Caucasus. Ethnobotany Research and Applications, 29, 1-21. ISI IF = 2.23
• Jolokhava T., Abdaladze O., Bakhia A., Ekhvaia J., Kikvidze Z.(2024): Soil pH as a strong driver of plant species distribution in alpine-nival ecotone of the Central Caucasus. Polish Journal of Soil Science (in press). ISI Impact Factor = 0.151 Manuscript URL: https://journals.umcs.pl/pjss/author/submission/16370    10.2023
• Jorjadze A., Kupradze I., Batsatsashvili K., R.W. (2024): Ethnobotany of the Caucasus. In: Bussmann, R.W. (eds) European Ethnobotany. Springer, Cham.
• Filipović , Glynis J., Kirleis W., Bogaard A., Ballantyne R., Charles M., Mosulishvili M., Mueller-Bieniek A., Padgett M., … Valamoti S-M. (202): Triticum timopheevii  s.l. (‘new glume wheat’) finds in regions of southern and eastern Europe across space and time. Vegetation History and Archaeobotany. 32, 6: 1-14. https://link.springer.com/article/10.1007/s00334-023-00954-w ISI  Impact factor = 2.55
• Dengler J, Jansen F, Chusova O, Hüllbusch E, Nobis MP, Van Meerbeek K, Akhmanová I, Bruun HH, Chytrý M, Guarino R, Karrer G, ..., Batsatsashvili K, ..., Nakhutsrishvili Get al. (2023): Ecological Indicator Values for Europe (EIVE) 1.0. Vegetation Classification and Survey 4: 7-29. http://doi.org/10.3897/VCS.98324 ISI Impact Factor = 1.510
• Novák P, Kalníková V, Szokala D, Aleksanyan A, Batsatsashvili K, Fayvush G, Kolbaia S, Nakhutsrishvili G, Sedláček V, Štěrba T, Zukal D. (2023): Transcaucasian Vegetation Database – a phytosociological database of the Southern Caucasus. Vegetation Classification and Survey4: 231 240. https://doi.org/10.3897/VCS.105521 ISI Impact Factor = 1.510
• Jorjadze A, Bussmann RW, Zambrana NY, Batsatsashvili K, Svanidze E.(2023): Traditional use of Polypores in Georgia (the Caucasus). Ethnobotany Research and Applications 26: 1-8. http://dx.doi.org/10.32859/era.26.5.1-18  ISI Impact Factor = 0.268
• Mumladze, L., Shetekauri, Sh., Barnaveli, N., Chelidze, D., Asanidze, Z. (2023): Species elevational richness gradient and species-area relationship in mountain vegetation of Javakheti highland (Georgia). Caucasiana, 2, pp.127-135.
• Asanidze, Z., Arabuli, T., Maghradze, E., Shavadze, L., Gogshelidze, M., Modebadze, N., Kiria, Barnaveli, N., Barjadze, Sh. (2023): Diversity and Ecology of the Soil- and Litter-dwelling Invertebrates and the Plant Associations in the Habitats of the Saskhori Limestone Quarry and Adjacent Areas, Eastern Georgia, Caucasus. Acta Zool. Bulg., 75 (3): 315-330. ISI Impact Factor = 0.362
• Lembrechts, J. J., van den Hoogen, J., Aalto, J., Ashcroft, M. B., De Frenne, P., Kemppinen, J., Kopecký, M., Luoto, M., Maclean, I. M. D., Crowther, T. W., Bailey, J. J., Haesen, S., Klinges, D. H., Niittynen, P., Scheffers, B. R., Van Meerbeek, K., Aartsma, P., Abdalaze, O., Abedi, M., … Lenoir, J. (2022): Global maps of soil temperature. Global Change Biology, 00, 1–35. https://doi.org/10.1111/gcb.16060 ISI Impact factor – 86
• Ekhvaia J., Bakhia A., Asanidze Z., Beltadze T., Abdaladze O. (2022): Leaf Functional Traits with Plant Resource Utilization Strategy in an Evergreen Scrub Species Rhododendron caucasicum Along Longitudinal Gradient in Georgia (The South Caucasus). Journal of Forest and Environmental Science Vol. 38, No. 2, pp. 110-121. https://doi.org/10.7747/JFES.2022.38.2.110 110 ISI Impact factor – 2.047
• McAlvay A., DiPaola,   D’Andrea C.,   Ruelle M. L.,   Mosulishvili M., Halstead P., Power A. G. (2022): Cereal Species Mixtures: An Ancient Practice with Potential for Climate Resilience. A Review.  Agronomy for Sustainable Development, 42 (5), 1-17. ISI Impact factor -- 5.832
• Inashvili T., Kupradze I., Batsatsashvili K. (2022): A Revised Catalog of Lichens of Georgia (South Caucasus)." Acta Mycologica57: 571 (2022), http//org/10.5586/am.57. ISI Impact factor -- 0.71
• Akobia, I., Janiashvili Z., Metreveli V., Zazanashvili N., Batsatsashvili K., Ugrekhelidze K. (2022): Modelling the Potential Distribution of Subalpine Birches (Betula ) in the Caucasus. Community Ecology 23, no. 2 (2022): 209-218. https://doi.org/10.1007/s42974-022-00097-4. ISI Impact factor – 1.66
• Kvlividze, Ekhvaia, J., Csiba, L., Asanidze, Z. (2022): Morphological and Genetic Research of Blackthorn (Prunus spinosa L.) in Georgia. The International Newsletter of the Partners of the Millennium Seed Bank Partnership (MSBP), Royal Botanic Gardens Kew, UK. Issue 3.
• Jorjadze, Batsatsashvili K., Kupradze I., Tigishvili K. (2022): Herbarium Catalogue of Georgian Fungi - Macromycetes. Tbilisi: Ilia State University.
• Kupradze I., Batsatsashvili K., Jorjadze A., Tigishvili K. (2022): Herbarium Catalogue of Georgian Lichens. Tbilisi: Ilia State University.
• Bürli S., Theurillat J-P., Winkler M., Lamprecht A., Pauli H., Rixen Ch., Steinbauer K., Wipf S., Abdaladze O., Andrews CH., Barančok P., Benito-Alonso J. L., Fernández Calzado M. R., Carranza M. L., Dick J., Erschbamer B., Ghosn D., Gigauri Kh., Kazakis G., Mallaun M., Michelsen O., Moiseev D., Moiseev P., Molau U., Molero Mesa J., Morra di Cella U., Nadeem I., Nagy L., Nicklas L., Palaj A., Pedersen B., Petey M., Puşcaş M., Rossi G., Stanisci A., Tomaselli M., Unterluggauer P., Ursu T-M., Villar L. & Vittoz P. (2021): Temperatures and growing season length define the upper limits of grasslands in European mountain. Alpine Botany. https://doi.org/10.1007/s00035-021-00250-1 ISI IF – 2.810
• Novák P., Zukal, Harásek M., Vlčková P., Abdaladze O., Willner W. (2021): Ecology and syntaxonomy of oak-hornbeam and ravine forests of the Eastern Greater Caucasus in Georgia. Folia Geobot https://doi.org/10.1007/s12224-020-09386-0 https://rdcu.be/cg7pv  ISI IF – 1.242  
• Abdaladze O., Zazanashvili N. (2021): Scientific Works of Professor Giorgi (Gia) Nakhutsrishvili. Bocconea 29: 9-19 https://doi.org/10.7320/Bocc29.009.
• Gigauri Kh., Abdaladze O., Bakhia A., Asanidze Z., Mamedova A. (2021): The first results of the 3rd cycle of Global Monitoring GLORIA Network of the Central Great Caucasus. Bocconea 29: 103-119 https://doi.org/10.7320/Bocc29.103.
• Nakhutsrishvili G., Abdaladze O., Batsatsashvili K. (2021): Ecological Gradients (West-East) and Vegetation of the Central Great Caucasus. Bocconea 29: 157-168 https://doi.org/10.7320/Bocc29.157.
• Tephnadze-Hoernchen N., Kikvidze Z., Nakhutsrishvili G., Abdaladze O. (2021): Subalpine vegetation along the soil moisture gradient under the climate change conditions: re-visitation approach (the Central Great Caucasus). Bocconea 29: 297-310 https://doi.org/10.7320/Bocc29.297.
• Jorjadze, A., K. Tigishvili, I. Kupradze, K. Batsatsashvili. (2021): "New species of gasteroid fungi for Georgia’s mycobiota." Annals of Agrarian Science19 (2021): 68-76.
• Bussmann, R. W., N.Y. Paniagua Zambrana, S. Sikharulidze, Z. Kikvidze, D. Kikodze, D. Tchelidze, K. Batsatsashvili, R. E. Hart. (2021): "An ethnobotany of Upper Imereti, Ukana Pshavi, Meshketi and Pankisi gorge, Sakartvelo (Republic of Georgia), Caucasus." Bocconea 29 (2021): 33-54.
• Goginashvili, N., Togonidze, N., Tvauri, I., Manvelidze, Z., Memiadze, N., Zerbe, S. and Asanidze, Z. (2021): Diversity and degradation of the vegetation of mountain belt forests of central Adjara (the Lesser Caucasus), Georgia. Journal of Forest Science, 67 (5), pp. 219-241. ISI IF = 0.89
• Mosulishvili M. (2021): Chapter 2. Endemic Wheat and Ancient Grains in Georgia.In: Cooper (ed.) Ancient Grains in Modern Soils.  DEStech Publications, PA, USA: 39-61.
• Mosulishvili M., Bedoshvili D., Maisaia I., Chkhutiashvili G., Ustiashvili  (2021): Aegilops tauschii, the D-genome donor of Triticum and the geographic origin of hexaploid wheat. Annals of Agrarian Science 18, 4: 437- 447.
• Mosulishvili M., Bedoshvili D., Maisaia I., Rusishvili N., Ustiashvili N., Chkhutiashvili G., Merabishvili (2021): Diversity of wheat in modern Georgia versus discoveries of the archaeological excavations of the 6.000 BC Neolithic sites of Lower Kartli Region (SE Georgia). Annals of Agrarian Science. 19,1: 46-52.
• Bedoshvili D., Mosulishvili M., Chkhutiashvili G., Maisaia I., Ustiashvili N., Merabishvili (2021): Diversity and local use of wheat in Georgia. Annals of Agrarian Science 19, 2: 103-110.
• Kazanci C., Oruç S., Mosulishvili M, L. Luczaj (2021): Wild Plants Used as Vegetables by Transhumant People Around the Georgia-Turkey Border in the Western Lesser Caucasus. Acta Societatis Botanicorum Poloniae 90 (906):1-23 DOI: 5586/asbp.906 ISI IF = 1.213
• Kazanci C., Oruç S., M. Mosulishvili (2021): Ethnoveterinary and fodder plants used among transhumant communities around the Georgia-Turkey border, in the Western Lesser Caucasus. Ethnobotany Research and Applications. 21, 28:1-26. ISI IF = 1.03
• Askerov E., Mallon D.P., Zazanashvili N., Kochishvili V., Sarukhanova S. (2021): An attempt to restore the population of the Goitered Gazelle (Gazella subgutturosa) in its indigenous range in the Caucasus (Mammalia: Bovidae). Zoology in the Middle East, 67 (3), pp. 189-197. ISI IF = 1.024
• Askerov E., Trepet S.A., Eskina T.G., Bibina K., Narkevich A., Pkhitikov A.B., Zazanashvili N., Akhmadova K. (2021): Estimation of the population densities of species prey or competitors to the leopard (Pantera pardus) in the Hyrcan National Park, Azerbaijan. Zoologichesky Jurnal, 100(8), pp. 947-955. ISI IF = 0.142
• სადუნიშვილი თ., მაისაია ი., ბაცაცაშვილი ქ., სიხარულიძე შ., დარჩიძე თ. (2021): საქართველოს აგრარული კულტურა. საქ. მეცნ. აკად. გამომც., თბილისი.

2006-2020 წწ.

რეფერირებად საერთაშორისო პერიოდულ გამოცემებში

1. Jolokhava T., Abdaladze O., Gadilia Sh., Kikvidze Z. (2020): Variable soil pH can drive changes in slope aspect preference of plants in alpine desert of the Central Great Caucasus (Kazbegi district, Georgia). Acta Oecologica 105, 103582 ISI
2. Kikvidze Z., Jolokhava T., Bakhia A., Abdaladze O. (2020): Jumping the barrier: does a glacier tongue affect species distribution along the elevation gradient in the subnival and nival belts? A case study of Mt. Kazbegi, Georgia, in the Central Great Caucasus Mountains. Botanica Serbica 44(2): 219-229. 
3. Jolokhava T., Abdaladze O., Asanidze Z., Kikvidze Z. (2020): Distribution of plant species and the rock particle size in subnival habitats of the Central Great Caucasus. Soil & Environment. 39(2): 194-203
4. Buchner, J., Yin, H., Frantz, D., Kuemmerle, T., Askerov, E., Bakuradze, T., Bleyhl, B., Elizbarashvili, N., Komarova, A., Lewińskaa, K.E., Rizayeva, A., Sayadyan, H., Bin Tank, B., Tepanosyan, G., Zazanashvili, N., Radelof, V.C. (2020): Land-cover change in the Caucasus Mountains since 1987 based on the topographic correction of multi-temporal Landsat composites. Remote Sensing of Environment, 248:111967
5. Bussman R. W., Zambrana N. P., Sikharulidze Sh., Kikvidze Z., Darchidze M., Ekhvaia J. et al. (2020): From the sea to the mountains: plant use in Ajara, Samegrelo and Kvemo Kartli, Sakartvelo (Republic of Georgia), Caucasus. Ethnobotany Research and Applications, 20 (9).
6. Murtazaliev R., Anatov Dzh., Ekhvaia J., Guseinova Z., Batsatsashvili K. (2020): Intraspecific variability of some functional traits of Trigonocaryum involucratum (Steen) Medw., a Caucasus endemic plant. Botanica Serbica, 44(2): 129-136. 
7. Mosulishvili M., Bedoshvili D., Maisaia I., Chkhutiashvili G. & N. Ustiashvili (2020): Aegilops tauschii, the D-genome donor of Triticum and the geographic origin of hexaploid wheat. Annals of Agrarian Science, 18, 4
8. Bedoshvili D., Mosulishvili M., Chkhutiashvili G., Chokheli M., Maisaia I. & N. Ustiashvili (2020):  Heritage wheats of Georgia. Annals of Agrarian Science, 18, 2: 123 - 129
9. Mosulishvili M., Bedoshvili D., Maisaia I., Rusishvili N., Chkhutiashvili G. & M. Merabishvili (2020): Diversity of wheat in modern Georgia versus discoveries of the archaeological excavations of the 6^th millennium BC Neolithic sites of Lower Kartli region. Annals of Agrarian Science 19, 1 
10. Kazancı C., Oruç S., & M. Mosulishvili (2020): Medicinal ethnobotany of wild plants: A cross-cultural comparison around Georgia-Turkey border, The Western Lesser Caucasus. Journal of Ethnobiology and Ethnomedicine
11. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K. and Hart, R.E. (2020): An ethnobotany of Kakheti and Kvemo Kartli, Sakartvelo (Republic of Georgia), Caucasus. Ethnobotany Research and Applications, 19: 1-27. 
12. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Darchidze, M., Manvelidze, Z., Ekhvaia, J., Kikodze, D., Khutsishvili, M., Batsatsashvili, K. and Hart, R.E. (2020): From the sea to the mountains-plant use in Adjara, Samegrelo and Kvemo Svaneti, Sakartvelo (Republic of Georgia), Caucasus. Ethnobotany Research and Applications, 20: 1-34. 
13. Goginashvili N., Ekhvaia J., Dobordjginidze R., Bachilava M., Tvauri I., Kobakhidze N. (2020): Comparative analysis of the three Caucasian oak taxa in Georgia (South Caucasus) based on leaf morphological variation. Annals of Agrarian Sciences, 18(8)
14. Kuemmerle, T., Bluhm, H., Ghoddousi, A., Arakelyan, M., Askerov, E., Bleyhl, B., Ghasabyan, M., Gavashelishvili, A., Heidelberg, A., Malkhasyan, A., Manvelyan, K., Soofi, M., Yarovenko, Yu., Weinberg, P. & Zazanashvili, N. (2020): Identifying priority areas for restoring mountain ungulates in the Caucasus ecoregion. Conservation Science and Practice. 276
15. Silakadze N., Kilian N., Korotkova N., Mosulishvili M. & T. Borsch. (2019): Multiple evolution of high mountain bellflowers with solitary flowers and calyx scales only renders a core of Campanula subg. Scapiflorae as a Caucasian clade. Systematics and Biodiversity, Volume 16, 4: 2-67 
16. Pistrick K., Akhalkatsi M., Ekhvaia J. (2019): Plant genetic resources of higher altitudes in southwestern Georgia (Samtskhe-Javakheti and Ajara). Genetics Resources and Crop Evolution. 
17. Bleyhl B., Arakelyan M., Askerov E., Bluhm H., Gavashelishvili A., Ghoddousi A., Heidelberg A., Khorozyan I., Malkhasyan A., Manvelyan K., Masoud M., Moqanaki E., Radeloff V.C., Soofi M., Weinberg P., Zazanashvili N. and Kümmerle T. (2019): Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Diversity and Distributions 25: 129-141
18. Askerov, E., Talibov, T., Manvelyan, K., Zazanashvili, N., Fatullayev, P. & Malkhasyan, A. (2019): Leopard (Panthera pardus) reoccupying its historic range in the South Caucasus: a first evidence (Mammalia: Felidae). Zoology in the Middle East 1-2: 88-90
19. Mosulishvili, M., D. Bedoshvili, I. Maisaia, G. Chkhutiashvili (2019): Georgia, the South Caucasus as the homeland of the hexaploid wheat. Annals of Agrarian Science. 17, 3: 287- 297
20. Zenderland J., Hart R., Bussmann R. W., Paniagua Zambrana N. Y., Sikharulidze Sh., Kikvidze Z., Kikodze D., Tchelidze D., Khutsishvili M., Batsatsashvili K. (2019): The Use of “Use Value”: Quantifying Importance in Ethnobotany. Economic Botany. 1-11

21. Kazanci C., Oruç S., Mosulishvili M. (2019): Cultural Keystone Species Without Boundaries: A Case Study on Wild Woody Plants of Transhumant People Around the Georgia-Turkey Border (the Western Lesser Caucasus). Journal of Ethnobiology. 39, 4: 1-30 

22. Ekhvaia J., Simeone M. C., Silakadze N., Abdaladze O. (2018): Morphological diversity and phylogeography of the Georgian durmast oak (Q. petraea subsp. iberica) and related Caucasian oak species in Georgia (South Caucasus). Tree Genetics & Genomes 14 (2), 17
23. Martín-Benito D., Pederson N., Köse N., Doğan M., Bugmann H., Mosulishvili M. & Ch. Bigler. (2018): Pervasive effects of drought on tree growth across a wide climatic gradient in the temperate forests of the Caucasus. Global Ecology and Biogeography 27: 1314-1325
24. Bussmann R. W., Paniagua Zambrana N. Y., Sikharulidze Sh., Kikvidze Z., Kikodze D., Tchelidze D., Batsatsashvili K., Hart R. E., Robbie E. (2018): Unequal brothers-Plant and fungal use in Guria and Racha, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 17, no. 1 (2018): 7-33
25. Kupradze I., Inashvili T., Batsatsashvili K., Lachashvili N., Gabelashvili S. (2018): Lichens of the arid region of David Gareji, Georgia (South Caucasus). Herzogia 31, no. 1: 268-275
26. Brander, L.M., Tankha, S., Sovann, G., Sanadiradze, G., Zazanashvili, N., Kharazishvili, D., Memiadze, N., Osepashvili, I., Beruchashvili, G., Arobelidze, N. (2018): Mapping the economic value of landslide regulation by forests. Ecosystem Services, vol. 32, part A., August 2018, pp. 101-109. 
27. Bleyhl, B., Arakelyan, M., Askerov, E., Bluhm, H., Gavashelishvili, A., Ghasabian, M., Ghoddousi, A., Hedelberg, A., Khorozyan, I., Malkhasyan, A., Manvelyan, K., Masoud, M., Moqanaki, E.M., Radeloff, V.C., Soofi, M., Weinberg, P., Zazanashvili, N. and Kuemmerle, T. (2018): Assessing niche overlap between domestic and threatened wild sheep to identify conservation priority areas. Biodiversity and Distribution. First published: 25 September 2018 
28. Nakhutsrishvili G. & Abdaladze O. (2017): Vegetation of the Central Great Caucasus along W-E and N-S Transects. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 11-16.
29. Nakhutsrishvili G. & Abdaladze O. (2017): Plant Diversity of the Central Great Caucasus. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 17-112.
30. Nakhutsrishvili, G. & Batsatsashvili, K. (2017): Quantitative analysis of the phytosociological relevés from the Central Greater Caucasus. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 113-132.
31. Randin Ch. F., Pellissier L., Guisan A., Nakhutsrishvili G. (2017): A Comparison of Climatic Niches of the Same Alpine Plant Species in the Central Caucasus and the Alps. In: Nakhutsrishvili, G., Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment. Springer, pp. 133-144.
32. Nakhutsrishvili, G., Batsatsashvili, K., Rudmann-Maurer, K., Körner, Ch. & Spehn, E. (2017): New Indicator values for Central Caucasus flora. In: Nakhutsrishvili, G. Abdaladze O., Batsatsashvili K., Körner Ch., Spehn E. (eds.) Plant Diversity in the Central Great Caucasus: A Quantitative Assessment, Springer, pp. 145-160.
33. Bussmann, R.W., Batsatsashvili, K., Kikvidze, Z., Khutsishvili, M., Maisaia, I., Sikharulidze, Sh., Tchelidze, D., Fayvush, G., Alexanyan, A., Mehdiyeva, N.P., Alizade, V.M., Paniagua Zambrana, N. Y. 2017. Ethnobotany of the Caucasus. Bussmann, R.W. (ed.). Springer. 708 p. 
34. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K., Hart, R.E. and Pieroni, A. (2017): Your poison in my pie - the use of Potato (Solanum tuberosum L.) leaves in Sakartvelo, Republic of Georgia, Caucasus, and Gollobordo, Eastern Albania. Economic Botany 
35. Mosulishvili M., Maisaia I., Batsatsashvili K. (2017): Proposal to conserve the name Triticum palaeocolchicum against T. karamyschevii (Poaceae). Taxon, 66, 12: 519-521 
36. Mumladze, L., Asanidze, Z., Walther, F. and Hausdorf, B. (2017): Beyond elevation: testing the climatic variability hypothesis vs Rapoport’s rule in vascular plant and snail species in the Caucasus. Biological Journal of the Linnean Society, p.blx027 
37. Mumladze, L., Ulrich, W., Asanidze, Z. and Japoshvili, G. (2017): An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Écoscience, pp.1-5
38. Mosulishvili M., Bedoschvili D., I. Maisaia. (2017): A consolidated list of Triticum species and varieties of Georgia to promote the repatriation of local diversity from foreign genebanks. Annals of Agrarian Science. 15, 1: 61-70.
39. Bakuradze T., Kandaurov A., Mosulishvili M., Nikolaishvili D., Gvilava M. & S. Kenkebashvili. (2017): Spatial Analysis Used in Baseline Study for the Preparation of Management Plan of Kazbegi Protected Areas. Earth Sciences. Special Issue: New Challenge for Geography: Landscape Dimensions of Sustainable Development. Vol. 6, No. 5-1, pp. 93-110 
40. Bleyhl, B., Baumann, M., Griffiths, P., Heidelberg, A., Manvelyan, K., Radeloff, V., Zazanashvili, N., Kuemmerle, T. (2017): Assessing landscape connectivity for large mammals in the Caucasus using Landsat 8 seasonal image composites. Remote Sensing of Environment, vol. 193:193-203
41. Breitenmoser, U., Askerov, E., Soofi, M., Breitemoser-Wursten, Heidelberg, A., Manvelian, K., and Zazanashvili, N. (2017): Transboundary leopard conservation in the Lesser Caucasus and Alborz Range. Cat News 65, pp. 24-25     http://fungi.biodiversity-georgia.net/ 
42. Winkler M., A. Lamprech, K. Steinbauer, K. Hülber, J-P. Treurillat, F. Breiner, P. Choler, S. Ertl, G.A. Giron, G. Rossi, P. Vittoz, M. Akhalkatsi, Ch. Bay, J.L. Alonso, T. Bergström, M.L. Carranza, E. Corcket, J. Dick, B. Ershbamer, R. F. Calzado, A. M. Forsaa, R. G. Gavilan, D. Chosn, Kh. Gigauri, D. Huber, R. Kanka, G. Kazakiz, M. Klipp. J. Kollar, T. Kudernatsch, P. Lersson, M. Mallaun, O. Michelsen, P. Moiseev, D. Moiseev, Ulf Mollau, J.M. Mesa, U.M. di Cella, L. Nagy, M. Petey, M. Puscas, Ch. Rixen, A., Stanisci, M., Suen, A. O., Syverhuset, M., Tomaselli, P., Unterluggauer, T., Ursu, L., Villar, M., Gottfried and Pauli, H. (2016): The rich sides of mountain summits-a pan-European view on aspect preferences of alpine plants. Journal of Biogeography 43: 2261-2273. 
43. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. (2016): Alpine plant distribution and thermic vegetation indicator on GLORIA summits in the Central Greater Caucasus. Pakistan Journal of Botany, 48(5): 1893-1902 
44. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Khutsishvili, M., Batsatsashvili, K. & Hart, R.E. (2016): A comparative ethnobotany of Khevsureti, Samtskhe-Javakheti, Tusheti, Svaneti, and Racha-Lechkhumi, Republic of Georgia (Sakartvelo), Caucasus. Journal of Ethnobiology and Ethnomedicine, 12(1), p.43.
45. The impacts of management and site conditions on the phytodiversity of the upper montane and subalpine belts in the Central Greater Caucasus. Tephnadze N., Abdaladze O., Nakhutsrishvili G., Simmering D., Waldhardt R., Otte A. Phytocoenologia, vol. 44, #3-4, pp. 255-291(37). 
46. Genetic relationships between wild progenitor pear (Pyrus L.) species and local cultivars native to Georgia, South Caucasus. Asanidze, Z., Akhalkatsi, M., Henk, A.D., Richards, C. M., Volk, G. M., Flora, 209, pp. 504-512. 2014
47. Tephnadze N., Abdaladze O., Nakhutsrishvili G., Simmering D., Waldhardt R., Otte A. (2013): Estimating the relative importance of land management and abiotic site conditions for phytodiversity in the Kazbegi district, Georgia. - Phytocoenosis
48. Cavieres, L., Pugnaire, F., Schöb, C., Xiao, S., Anthelme, F., Björk, R., Dickinson, K., Cranston, B., Gavilán, R., Gutiérrez-Girón, A., Kanka, R., Kikvidze Z., Maalouf, J.-P., Mark, A., Noroozi, J., Parajuli, R., Phoenix, G., Reid, A., Ridenour, W., Rixen, C., Wipf, S., Zhao, L., Escudero, A., Zaitchik, B., Aschehoug, E., Lingua, E. and Callaway, R. (2013): Facilitative plant interactions and climate simultaneously drive alpine plant diversity. – Ecology Letters 
49. Jimbo, K., Kubota, K. & Kikvidze, Z. (2013): Ground beetle succession on Mount Fujisan. – Biogeography (Japan). 15
50. Asensio, C., Lozano, F.J., Ortega, E., & Kikvidze, Z. (2013): Study on the effectiveness of an agricultural technique based on aeolian deposition, in a semiarid environment. Environmental Engineering and Management Journal 12
51. Ekhvaia J., Gurushidze M, Blattner F.R., Akhalkatsi M. (2013): Genetic diversity of Vitis vinifera in Georgia: relationships between local cultivars and wild grapevine, V. vinifera subsp. sylvestris. - Theoretical and Applied Genetics 
52. Kürschner, H., Batsatsashvili, K. & Parolly, G. 2013. Noteworthy additions to the bryophyte flora of Georgia. – Herzogia 26: 213 –216.
53. Montalvo Mancheno, C.S., Nakhutsrishvili, G. & Zazanashvili, N. (2013): A Perspective on Ecological Corridor for Maintaining Healthy Ecological Processes in the Caucasus. Research in Protected Areas, Mittersill, pp. 523-526.
54. Barjadze, Sh., Asanidze, Z. (2013): Morphometric investigation of grapevine and willow root-feeding aphid, Pemphigus saliciradicis (Börner, 1950) (Hemiptera: Aphididae: Eriosomatinae). - Dtsch. Entomol. Z. 60 (1), 45–52 
55. Butterfield, B.J., L. A. Cavieres, R. M. Callaway, B. J. Cook, Z. Kikvidze, C. J. Lortie, R. Michalet, F. I. Pugnaire, C. Sch€ob, S. Xiao, B. Zaitchek, F. Anthelme, R. G. Bj€ork, K. Dickinson, R. Gavilan, R. Kanka, J.-P. Maalouf, J. Noroozi, R. Parajuli, G. K. Phoenix, A. Reid, W. Ridenour, C. Rixen, S. Wipf, L. Zhao, R. W. Brooker (2013): Alpine cushion plants inhibit the loss of phylogenetic diversity in severe environments. - Ecology Letters, 1-8. 
56. Miyata M., Wu B., Kikvidze Z., Fukuda K. (2013): Mycorrhizal Cirsium purpuratum improves the growth of Clematis stans in volcanic scoria of Mount Fuji. - Micoscience, vol. 54, (4): 279-284.
57. Khojasteh F., M. A. Z. Chahouki, H. Azarnivand, Z. Kikvidze (2013): Life form and preference can drive spatial relationships among plant species in semi-arid rangelands of middle Iran. - The Rangeland Journal. 
58. Suzuki M, Miyashita T, Kabaya H, Ochiai K, Asada M, & Kikvidze Z. (2012): Deer herbivory as an important driver of divergence of ground vegetation communities in temperate forests. - Oikos. 
59.  Abukava K, Yamamuro M, Kikvidze Z, Asada A, Xu C, & Sugimoto K. (2012): Assessing the biomass and distribution of submerged aquatic vegetation using multibeam echo sounding in Lake Towada, Japan. - Limnology
60. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. (2013): Monitoring of vascular plant diversity in the alpine zone of the Central Greater Caucasus in a changing climate. Turkish Journal of Botany, 37: 1104-1114. 
61. Pauli H., Gottfried M., Dullinger S., Abdaladze O., Akhalkatsi M., Benito Alonso J. L., Coldea G., Dick J., Erschbamer B., Calzado R. F., Ghosn D., Holten J. I., Kanka R., Kazakis G., Kollár J., Larsson P., Moiseev P., Moiseev D., Molau U., Mesa J. M., Nagy L., Pelino G., Puşcaş M., Rossi G., Stanisci A., Syverhuset A. O., Theurillat J-P., Tomaselli M., Unterluggauer P., Villar L., Vittoz P., Grabherr G. (2012): Recent plant diversity changes on Europe’s mountain summits. Science vol. 336, pp. 353-355. 
62. Gottfried, M., Pauli, H., Futschik, A., Akhalkatsi, M., Barančok P., Alonso, J.L.B., Coldea, G., Dick, J., Erschbamer, B., Rosa, M., Calzado, F., Kazakis G., Krajči, J., Larsson, P., Mallaun, M., Michelsen, O., Moiseev, D.15, Moiseev, P., Molau, U., Merzouki, A., Nagy, L., Nakhutsrishvili, G., Pedersen, B., Pelino, G., Puscas, M., Rossi, G., Stanisci, A., Theurillat, J.-P., Tomaselli, M., Villar, L., Vittoz, P., Vogiatzakis, I., Grabherr G. (2012): An early warning signal for a continental-scale response to ongoing climate change. Nature Climate Change 2, 111–115.
63. Caprez, R., Spehn, E., Nakhutsrishvili, G., Körner, Ch. (2011): Drought at erosion edges selects for a ‘hidden’ keystone species. Plant Ecology & Diversity. ISSN: 1755-0874. 
64. Asanidze, Z., Akhalkatsi, M., Gvritishvili, M. (2011): Comparative morphometric study and relationships between the Caucasian species of wild pear (Pyrus spp.) and local cultivars in Georgia. Flora 206, 11
65. Nakhutsrishvili G. (2011): La végétation des zones de haute montagne du Caucase. Ethnopharmacologia. Nr. 47: 45-51. 
66. Nakhutsrishvili G. (2011): Historique de l'Institut de Botanique de Géorgie. Ethnopharmacologia. Nr. 47: 39-44. 
67. Akhalkatsi M., Ekhvaia, J., Mosulishvili M., Nakhutsrishvili G., Abdaladze O., Batsatsashvili K. (2010): Reasons and processes leading to the erosion of crop genetic diversity in mountainous regions of Georgia. Mountain Research and Development. 30, 3:304-310. 
68. Ekhvaia J., Akhalkatsi M. (2010): Morphological variation and relationships of Georgian populations of Vitis vinifera L. subsp. sylvestris (C.C. Gmel.) Hegi. Flora 205, 9:608-617.
69. Inashvili, Tz. & Batsatsashvili, K. (2010): New Lichen Records from Georgia. Turkish Journal of Botany. 34(6): 549-553. 
70. Erschbammer B., Mallaun M., Unterluggauer P., Abdaladze O., Akhalkatsi M., Nakhutsrishvili G. (2010): Species richness and distribution above the treeline in the Central Alps (Italy) and in the Central Caucasus (Georgia). Tuexenia. 30:11-29. (ISSN: 0722-494X) 
71. Hughes N.M., Johnson D.M., Akhalkatsi M., Abdaladze O. (2009): Characterizing Betula litwinowii Seedling Microsites at the Alpine-Treeline Ecotone, Central Greater Caucasus Mountains, Georgia. – Arctic, Antarctic & Alpine Research, vol. 41, No 1, pp. 112-118. 
72. Dullinger S., I. Kleinbauer, H. Pauli, M. Gottfried, Ch. Klettner, R. Brooker, L. Nagy, J.-P. Theurillat, J. I. Holten, O. Abdaladze, J-L. Benito, J.-L. Borel, G. Coldea, D. Ghosn, R. Kanka, A. Merzouki, P. Moiseev, U. Molau, K. Reiter, G. Rossi, A. Stancisi, M. Tomaselli, P. Unterlugauer, and G. Grabherr (2007): Do co-occurrence patterns in alpine plant communities indicate a stress-driven increase in facilitation? - Journal of Ecology.  
73. Akhalkatsi M., Abdaladze O., Nakhutsrishvili G., Smith W. K. (2006): Rhododendron caucasicum and Microtopography Extend the Betula litwinowii Alpine Treeline (Caucasus Mountains, Georgia). – Arctic, Antarctic & Alpine Research, vol. 38, #4, pp. 441-448. ISSN: 1523-0430

სხვა პუბლიკაციები 

1. Asanidze, E., Asanidze, Z., Lordkipanidze, , Tsertsvadze, M., Asanidze, A., Kristesashvili, J., Jibladze, A., Chodnekar, S., Parunashvili, N., Asanidze, B. (2026): Work-Life Realities of Doctors in Georgia and Its Impact on Mental Health and Quality of Life. PLOS ONE. Status of the article: In press, Under review. ISI IF = 2.6
2. NakhutsriSvili G., Abdaladze O., Batsatsashvili K. (2026): High Mountain Plant Communities of the Central Great Caucasus. Yunnan, Kunming, China (in press).
3. Zazanashvili N., M. Garforth and G. Nakhutsrishvili (2021): Caucasus-Anatolian-Hyrcanian Temperate Forests. Chapter 50035 for the Encyclopedia of Conservation.  Elsevier
4. Jolokhava T., Kikvidze Z., Gigauri Kh., Abdaladze O. (2021): Maximum soil temperature, plant available NPK, pH and soil organic matter (SOM) along the altitudinal gradient (3000-3900 m a.s.l.) in the Central Great Caucasus. 
5. Burgaz A.R., Teuvo A., Inashvili T., Batsatsashvili K., Kupradze I. (2018): Study of Georgian Cladoniaceae. Botanica complutensis 42: 19-55
6. Maharramova, E., Moqanaki, E.M., Askerov, E., Faezi, S., Alinezhad, H., Mousavi, M., Kuemmerle, T., Heidelberg, A. & Zazanashvili, N. (2018): Transboundary leopard movement between Azerbaijan and Iran in the Southern Caucasus. Cat News 67: 8-10
7. Kuemmerle, T., Blehyl, B., Bluhm, H., Ghoddousi, A., Heidelberg, A., Sarukhanova, S., Zazanashvili, N. (2018): Identifying priority areas for restoring large mammal populations in the Caucasus Ecoregion. 5th European Congress of Conservation Biology, 12-15 June, Jyväscylä, Finland 
8. Gavrilović M., Tešević V., Đorđević I., Rajčević N., Bakhia A., Garcia Jacas N., Susanna A., Marin P. D., Janaćković P. (2018): Leaf micromorphology, antioxidative activity and a new record of 3-deoxyamphoricarpolide of relict and limestone endemic Amphoricarpos elegans Albov (Compositae) from Georgia. Archives of Biological Sciences 2018 OnLine-First Issue 00, Pages: 22-22
9. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2017): Ethnobotany of Samtskhe-Javakheti, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 16 (1):7-24 
10. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2017): Plants in the spa – the medicinal plant market of Borjomi, Sakartvelo (Republic of Georgia), Caucasus. Indian journal of traditional knowledge 16 (1): 25-34
11. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, Sh., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K., Hart, R.E. and Robbie, E. (2018): Unequal brothers–Plant and fungal use in Guria and Racha, Sakartvelo (Republic of Georgia), Caucasus. Indian Journal of Traditional Knowledge 17(1): 7-33
12. Bussmann, R.W., Zambrana, N.Y.P., Sikharulidze, S., Kikvidze, Z., Kikodze, D., Tchelidze, D., Batsatsashvili, K. & Hart, R.E. (2016): Medicinal and Food Plants of Svaneti and Lechkhumi, Sakartvelo (Republic of Georgia), Caucasus. Medicinal and Aromatic Plants (Los Angeles), 5: 266 
13. Abdaladze O., Nakhutsrishvili G., Batsatsashvili K., Gigauri Kh., Jolokhava T., Mikeladze G. (2015): Sensitive Alpine Plant Communities to the Global Environmental Changes (Kazbegi Region, the Central Great Caucasus). - American Journal of Environmental Protection. Special Issue. Applied Ecology: Problems, Innovations. Vol. 4, No.3-1, pp. 93-100.
14. Askerov E., Talibov T., Manvelyan K., Zazanashvili N., Malkhasyan A., Fatullayev P. & Heidelberg A. (2015): South-Eastern Lesser Caucasus: the most important landscape for conserving the Leopard (Panthera pardus) in the Caucasus region (Mammalia: Felidae). - Zoology in the Middle East,v. 61, 2.
15. Kupradze I., Jorjadze I., A. Arabidze, T. Beltadze, Batsatsashvili K. (2015): Ethnobiological Study of Svaneti Fungi and Lichens: History of Research, Diversity, Local Names and Traditional Use. – American Journ. of Environmental Protection. Special Issue. Applied Ecology: Problems, Innovations. Vol. 4, No. 3-1, pp. 101-110.
16. Akhalkatsi M., Asanidze Z. (2015): Pyrus caucasica Fed. Ancestor of Georgian Pear Landraces. – Science, technology and development, 34 92), pp. 105-108.
17. Wine, Beer, Snuff, Medicine, and Loss of Diversity: Ethnobotanical Travels in the Georgian Caucasus. Bussmann, R. W., Paniagua Zambrana N. Y., Sikharulidze Sh., Kikvidze Z., Kikodze D., Jinjikhadze T., Shanshiashvili T., Chelidze D., Batsatsashvili K. & Bakanidze N. Ethnobotany Research and Applications, 12: 237-313, 2014
18. Caucasus endemic vascular plant species Red List assessments (385 taxa). Scharz, G.E., Shulkina, T.V., Nakhutsrishvili, G., Batsatsashvili, K. et al. (evaluators). The IUCN Red List of Threatened Species, 2014
19. Vascular plant diversity and climate change in the alpine zone of the Central Greater Caucasus. Gigauri Kh., Akhalkatsi M., Abdaladze O., Nakhutsrishvili G. International Journal of Ecosystems and Ecology Science. ISSN 2224-4980, v. 4 (4), pp. 573-589, 2014
20. Waldhart R., Abdaladze O., Otte A., Simmering D. (2011): Ursachen und Folgen des Landschaftswandels im Kaukasus Georgiens: Interdisziplinäre Forschung für eine nachhaltigere Zukunft. Spiegel der Forschung. 28, Nr 2: 4-15
21. Otte, A., Akhaltkatsi, M., Nakhutsrishvili, G., Simmering, D., Waldhardt, R. (2011): Phytodiversität in Georgien: Die Bedeutung von Standort und Landnutzung im Großen und Kleinen Kaukasus. Spiegel der Forschung 28, Nr. 2:24-31
22. Pistrick K., Akhalkatsi M., Girgvliani T., Shanshiashvili T. (2010): Sammlung von Saat- und Pflanzgut pflanzengenetischer Ressourcen in Svanetien und Lečchumi (Georgien, Kaukasus) 2009. Berichte Ges. Pflanzenbauwiss. 5, 1:125-127
23. Mosulishvili M., Arabuli G. (2010): Buxus Stands in East Georgia. Bull. of Georg. Nation. Museum, #2, pp. 128-132. 
24. Lachashvili N., Khetsuriani L., Batsatsashvili K. (2010): Effect of grazing on some communities of Bothriochloa ischaemum in Shida Kartli (East Georgia). Bull. of Georgian National Academy of Sciences. 8, #1-2, pp. 20-25.
25. Nakhutsrishvili G., Akhalkati M., Abdaladze O. (2009): Main Threats to the Mountain Biodiversity in Georgia (the Caucasus). – Mountain Forum Bulletin, vol. IX, Issue 2, pp. 18-19.
26. Pistrick, K., Akhalkatsi, M., Girgvliani, T., Shanshiashvili, T. (2009): Collecting plant genetic resources in Upper Svaneti (Georgia, Caucasus mountains). J. Agric. Rural Dev. Trop. Subtrop. Supplement 92:127-135.
27. Schultzke, R., Zazanashvili, N. and Osepashvili, I. (2009): Restoration of forest landscapes in the Southern Caucasus. Pp. 119-125 in: Bodegom, V, Jan, A., Savenije, H. and Wit, M. (Eds.). Forest and Climate Change: adaptation and mitigation. Tropendos International, Wageningen, The Netherlands.
28. Batsatsashvili K., Chelidze N. (2009): Lichens as Bioindicators of Tbilisi Air Pollution. TSU Publ., pp. 295–309 (Georgian). 
29. Bohn U., Zazanashvili N., Nakhutsrishvili G. (2007): The Map of the Natural Vegetation of Europe and its application in the Caucasus Ecoregion. – Bulletin of the Georgian National Academy of Sciences, col. 175, #1, pp. 112-121.

სამეცნიერო კრებულებში:

1. Zazanashvili, N., Sanadiradze, G., Garforth, M., Bitsadze, M., Manvelyan, K., Askerov, E., Mousavi, M., Krever, V., Shmunk, V., Kalem, S. and Devranoğlu Tavsel, S., eds. (2020): Ecoregional Conservation Plan for the Caucasus: 2020 Edition. WWF, KfW, Tbilisi.
2. Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. (2020): Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
3. Zazanashvili, N., Manvelyan, K., Askerov, E., Mousavi, M., Krever, V., Kalem, S. & Garforth, M. (2020): The boundaries and bio-physical features of the Caucasus Ecoregion. Pp. 9-20 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
4. Bitsadze, M., Manvelyan, K., Askerov, E., Mousavi, M., Shmunk, V., Kalem, S., Zazanashvili, N., Mnatsekanov, R., Devranoglu Tavsel, S., Cevik, M., Ahmadova, K., Beruchashvili, G., Kandaryan, A. & Garforth, M. (2020): Key Biodiversity Areas in the Caucasus Ecoregion. Pp. 21- 28 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
5. Zazanashvili, N., Manvelyan, K., Askerov, E., Mousavi, M., Shmunk, V., Kalem, S., Beruchashvili, G., Bitsadze, M. & Garforth, M. (2020): Conservation landscapes in the Caucasus Ecoregion. Pp. 29 - 36 in Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
6. Askerov, E., Manvelyan, K., Gurielidze, Z., Mousavi, M., Shmunk, V., Trepet, S., Kütükçü, A. E., Heidelberg, A. & Zazanashvili, N. (2020): Status of large carnivores in the Caucasus. Pp. 37 - 47 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
7. Weinberg, P., Askerov, E., Manvelyan, K., Gurielidze, Z., Mousavi, M., Kütükçü, A.E., Heidelberg, A. & Zazanashvili, N. (2020): Status of large herbivores in the Caucasus. Pp. 48 - 71 in: Zazanashvili, N., Garforth, M. and Bitsadze, M., eds. Ecoregional Conservation Plan for the Caucasus, 2020 Edition: Supplementary Reports. WWF, KfW, Tbilisi.
8. Abdaladze O. (2011): Functional Diversity of Alpine Plants of Georgia and Climate Global Change. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Science of Georgia, Tbilisi: 15-18. 
9. Nakhutsrishvili G. (2011): Actual Problems of Biodiversity. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Sci. of Georgia, Tbilisi: 133-134. 
10. Gigauri Kh., Akhalkatsi M., Nakhutsrishvili G., Abdaladze O. (2011): First Signs of Weak Thermophilisation on the GLORIA-EUROPE Target Region in the Central Caucasus. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Sci. of Georgia, Tbilisi: 19-22.
11. Sharikadze Kh., Nakhutsrishvili G., Akhalkatsi M., Abdaladze O. (2011): Species Diversity of Eroded Slopes in the Kazbegi region. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Sci. of Georgia, Tbilisi: 29-32. (Georgian with English summary).
12. Zazanashvili N., Gavashelishvili A., Montalvo K., Beruchashvili G. (2011): Modelling the Response of South Caucasus Forest Classes to Climate Change. In: G. Nakhutsrishvili et al. (eds). Biodiversity of Georgia. Nat. Acad. of Sci. of Georgia, Tbilisi: 210-212.
13. Akhalkatsi M., Fritsch R. M., Nakhutsrishvili G., Pistrik K. (2008): Habitats of Allium species in Georgia.– In: M. Keusgen & M. Fritsch (eds.): Botany, Taxonomy and Phytochemistry of Wild Allium L. Species of the Caucasus and Central Asia. Marburg-Gatersleben, pp. 147-156
14. Pistrik K., Akhalkatsi M., Nakhutsrishvili G. (2008): Collecting wild Allium species in Georgia. – In: M. Keusgen & M. Fritsch (eds.): Botany, Taxonomy and Phytochemistry of Wild Allium L. Species of the Caucasus and Central Asia. Marburg-Gatersleben, pp. 45-52

წიგნები და თავები წიგნებში:

1. ნახუცრიშვილი გ., აბდალაძე ო., ბაცაცაშვილი ქ., ძაძამია ლ. (2023): საქართველოს ბუნებრივი ტყეები (სამხრეთ კავკასია).
2. Nakhutsrishvili G., Abdaladze O., Batsatsashvili K., Dzadzamia L. (2023): Natural Forests of Georgia (the South Caucasus)“.
3. Bussmann, R.W., Batsatsashvili, K.,  Kikvidze, Z.,  Paniagua Zambrana, N.Y., Khutsishvili, M., Maisaia, I., Sikharulidze, Sh., Tchelidze, D. et al. (2020): Ethnobotany of the Mountain Regions of Far Eastern Europe. Batsatsashvili, K., Kikvidze, Z., Bussmann, R.W. (eds.). Springer. 1063p.
4. Bussmann, R.W., Batsatsashvili, K.,  Kikvidze, Z. et al. (2020): Ethnobotany of the Mountain Regions of Central Asia and Altai. Batsatsashvili, K., Kikvidze, Z., Bussmann, R.W. (eds.). Springer. 881p.
5. Mosulishvili M., Bedoshvili D., Maisaia I., Rusishvili N., Chkhutiashvili G. (2019): The Ancient, Traditional Georgian Wheat Species and Their Role in Wheat Evolution. In: Proceedings of the International Conference “Wheat in European countries and Georgia as one of the centres of origin of Domesticated Wheat”: 232 – 239. Tbilisi, Georgia.
6. Gambashidze G., Jolokhava T. et al. (2019): Soil erosion modelling using RUSLE 1 in Shida Kartli Region, Georgia. Food and Agriculture Organization of the United Nations, Rome, Italy.
7. ნახუცრიშვილი გ., აბდალაძე ო. (2018): ბოტანიკური და ეკოლოგიური კვლევები ყაზბეგის რეგიონში. ნ. ანთიძე (რედ.) მუზეუმი და კულტურული მემკვიდრეობა. თბილისი, გვ. 205–210.
8. გოგინაშვილი, ნ., თვაური, ი., მანველიძე, ზ., მემიაძე, ნ., ასანიძე, ზ., ღამბაშიძე, გ. (2018): აჭარის ტყის იშვიათი მერქნიანი სახეობების ინვენტარიზაციის მასალები. საქართველოს გარემოსა და სოფლის მეურნეობის სამინისტროს სოფლის მეურნეობის სამეცნიერო კვლევითი ცენტრი, გამომც. „უნივერსალი“, თბილისი.
9. Nakhutsishvili G., Batsatsashvili K., Gagnidze R., Shetekauri Sh., Manvelidze Z., Memiadze N. & D. Kharazishvili (2013): Georgia. – In: Schatz, G., Shulkina, T. & Solomon, J. (eds.) Red List of the Endemic Plants of the Caucasus Region. Missouri Botanical Garden Press, Saint Louis, USA 
10. Nakhutsrishvili, G. (2013): The vegetation of Georgia (Caucasus). - Springer-Verlag, Heidelberg, New York, Dordrecht, London, 235 pp.
11. ნ. ელიზბარაშვილი, ნ.ზაზანაშვილი, დ. ნიკოლაიშვილი, ლ. მაჭავარიანი, გ. მელაძე, გ. დვალიშვილი. საქართველოს ბუნება. გამომც. „პალიტრა L.“, თბილისი, 294 გვ. 2013.
12. Nakhutsrishvili, G., Zazanashvili, N. & Batsatsashvili, K. (2011): Why the Colchis and Hyrcan relict forests of the western Eurasian Caucasus should be considered “temperate rainforest”. – In: DellaSala D. (ed.) Temperate and Boreal Rainforests of the World: Ecology and Conservation. Island Press, pp. 214-221.
13. Bondirev, I.V., Seperteladze, Z. KH., Zazanashvili, N.I., Tavartkiladze, A.M., Budagov, B.A., Museibov, M.A., Vardanian, T.G., Balian, A.S. (2009): Critical changes of the ecosystems in Southern Caucasus. Pp. 7-14 in I. V. Bondirev, A. M. Tavartkiladze (Eds.). Critical ecoregions: modern conditions, problems and ways of their decisions. Publishing House. Tbilisi: "Polygraph".
14. Nakhutsrishvili G., Abdaladze O., Akhalkatsi M. (2006): Biotope Types of the Treeline in the Central Greater Caucasus. – In: D. Gafta & J. Akereoyd (eds.): “Nature Conservation: Concepts and Practice”, Springer Verlag, pp. 211-225. ISBN 9783540472292.
15. Körner, Ch., Nakhutsrishvili, G., Spehn, E. M. (2006): High-Elevation Land Use, Biodiversity and Ecosystem Functioning. In: Spehn, E. M., Liberman, M., Kőrner, Ch. (Eds.), Land Use Change and Mountain Biodiversity, Taylor & Francis Group, Boca Raton, London, New York, pp. 3-21.
16. Nakhutsrishvili G., Sikharulidze Sh., Abdaladze O. (2006): Bakuriani – Natural and Cultural Resources of the Borjomi Region. – Institute of Botany, Tbilisi, 56 p.